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plantain des sables, sand plantain

hoary plantain, plantain moyen

Habit Annuals; roots taproots, slender. Perennials; caudex glabrous or hairy; roots taproots, thick.
Stems

100–350 mm, freely branched.

0–20 mm.

Leaves

cauline, opposite, 60–80 × 1–3 mm;

blade linear to linear-lanceolate, margins entire or toothed, veins conspicuous or not, surfaces hairy.

prostrate, sometimes ascending, 40–200 × 30–70 mm;

blade elliptic to ovate, margins entire or toothed, veins conspicuous, surfaces hairy, hairs 0.5 mm.

Scapes

100–650 mm, hairy.

80–300 mm, not groove-angled, hirsute.

Spikes

greenish or brownish, (50–)150–250 mm, densely flowered, eglandular;

bracts proximal strongly differing from distal, ovate, 2–5(–7) mm, length 1–1.5 times sepals, proximal bracts: apex acute.

grayish or whitish, 100–400 mm, densely flowered, shiny;

corolla lobes of neighboring flowers overlapping;

bracts ovate, 1.8–2 mm, length 0.9–1 times sepals.

Flowers

sepals 2–3 mm;

corolla radially symmetric, lobes reflexed, 2–4 mm, base obtuse;

stamens 4.

sepals 2 mm, adaxial 2 nearly distinct;

corolla radially symmetric, lobes reflexed, 1.5 mm, base obtuse;

stamens 4.

Seeds

2, 2–2.5 mm.

2–4, 2 mm.

2n

= 12.

= 12, 24.

Plantago indica

Plantago media

Phenology Flowering late summer–fall. Flowering summer.
Habitat Roadsides, railroads, sandy shorelines. Disturbed areas, neutral and basic soils.
Elevation 0–200 m. (0–700 ft.) 0–200 m. (0–700 ft.)
Distribution
from FNA
CA; CT; DE; IA; IL; IN; MA; ME; MI; MN; MO; NC; NH; NJ; NY; OH; OR; PA; VA; VT; WA; WI; BC; MB; ON; QC; Eurasia [Introduced in North America]
from FNA
CT; IL; MA; ME; MI; NH; NJ; NY; PA; RI; WI; MB; NB; ON; QC; Eurasia [Introduced in North America]
[BONAP county map]
Discussion

While considering the proposal made by W. L. Applequist (2006) to reject Plantago psyllium, the Nomenclature Committee for Vascular Plants also decided that P. indica is a legitimate name (R. K. Brummitt 2009). A. B. Doweld and A. Shipunov (2017) published a proposal to reject P. indica in favor of P. arenaria. That proposal awaits a decision by that committee.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Three varieties of Plantago media have been recognized in western Europe. It is not clear how names of those varieties apply to P. media in eastern Europe and North America.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 17, p. 288. FNA vol. 17, p. 290.
Parent taxa Plantaginaceae > Plantago Plantaginaceae > Plantago
Sibling taxa
P. afra, P. argyrea, P. aristata, P. australis, P. canescens, P. cordata, P. coronopus, P. elongata, P. erecta, P. eriopoda, P. firma, P. floccosa, P. helleri, P. heterophylla, P. hookeriana, P. lanceolata, P. macrocarpa, P. major, P. maritima, P. media, P. ovata, P. patagonica, P. pusilla, P. rhodosperma, P. rugelii, P. sempervirens, P. sparsiflora, P. subnuda, P. tweedyi, P. virginica, P. wrightiana
P. afra, P. argyrea, P. aristata, P. australis, P. canescens, P. cordata, P. coronopus, P. elongata, P. erecta, P. eriopoda, P. firma, P. floccosa, P. helleri, P. heterophylla, P. hookeriana, P. indica, P. lanceolata, P. macrocarpa, P. major, P. maritima, P. ovata, P. patagonica, P. pusilla, P. rhodosperma, P. rugelii, P. sempervirens, P. sparsiflora, P. subnuda, P. tweedyi, P. virginica, P. wrightiana
Synonyms P. arenaria, P. psyllium
Name authority Linnaeus: Syst. Nat., ed. 10, 2: 896. 1759, legitimacy of — name in question Linnaeus: Sp. Pl. 1: 113. (1753)
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