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plantain des sables, sand plantain

Mexican plantain

Habit Annuals; roots taproots, slender. Perennials; caudex glabrous; roots fibrous, stout.
Stems

100–350 mm, freely branched.

0–10 mm.

Leaves

cauline, opposite, 60–80 × 1–3 mm;

blade linear to linear-lanceolate, margins entire or toothed, veins conspicuous or not, surfaces hairy.

40–350 × 6–77 mm;

blade elliptic to narrowly elliptic, margins entire, veins conspicuous, surfaces pilose, rarely glabrate, adaxial surface hairs not floccose, less than 2 mm long, more than 0.03 mm wide.

Scapes

100–650 mm, hairy.

30–560 mm, hairy, hairs antrorse, short.

Spikes

greenish or brownish, (50–)150–250 mm, densely flowered, eglandular;

bracts proximal strongly differing from distal, ovate, 2–5(–7) mm, length 1–1.5 times sepals, proximal bracts: apex acute.

greenish or brownish, 100–1000 mm, densely flowered;

bracts narrowly triangular, 1.6–4.2 mm, length 0.8–1.5 times sepals.

Flowers

sepals 2–3 mm;

corolla radially symmetric, lobes reflexed, 2–4 mm, base obtuse;

stamens 4.

sepals 2–2.5 mm;

corolla radially symmetric, lobes erect, forming a beak, 2–2.8 mm, base obtuse;

stamens 4.

Seeds

2, 2–2.5 mm.

3, 1.2–2.2 mm.

2n

= 12.

= 24.

Plantago indica

Plantago australis

Phenology Flowering late summer–fall. Flowering summer.
Habitat Roadsides, railroads, sandy shorelines. Open places.
Elevation 0–200 m. (0–700 ft.) 0–1000 m. (0–3300 ft.)
Distribution
from FNA
CA; CT; DE; IA; IL; IN; MA; ME; MI; MN; MO; NC; NH; NJ; NY; OH; OR; PA; VA; VT; WA; WI; BC; MB; ON; QC; Eurasia [Introduced in North America]
from FNA
AZ; Mexico; Central America; South America
[WildflowerSearch map]
[BONAP county map]
Discussion

While considering the proposal made by W. L. Applequist (2006) to reject Plantago psyllium, the Nomenclature Committee for Vascular Plants also decided that P. indica is a legitimate name (R. K. Brummitt 2009). A. B. Doweld and A. Shipunov (2017) published a proposal to reject P. indica in favor of P. arenaria. That proposal awaits a decision by that committee.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Plantago australis occurs in Cochise, Coconino, and Pima counties. Plantago australis is most diverse in South America, where as many as 16 subspecies (K. Rahn 1974) may be recognized. Plants from California identified as P. hirtella are most likely P. subnuda. However, since the most important distinguishing character of P. australis is the absence of the developed taproot (which is fragile and often broken in herbarium specimens), all these samples require careful examination. Further research is needed also to clarify the circumscriptions of P. australis and P. subnuda.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 17, p. 288. FNA vol. 17, p. 284.
Parent taxa Plantaginaceae > Plantago Plantaginaceae > Plantago
Sibling taxa
P. afra, P. argyrea, P. aristata, P. australis, P. canescens, P. cordata, P. coronopus, P. elongata, P. erecta, P. eriopoda, P. firma, P. floccosa, P. helleri, P. heterophylla, P. hookeriana, P. lanceolata, P. macrocarpa, P. major, P. maritima, P. media, P. ovata, P. patagonica, P. pusilla, P. rhodosperma, P. rugelii, P. sempervirens, P. sparsiflora, P. subnuda, P. tweedyi, P. virginica, P. wrightiana
P. afra, P. argyrea, P. aristata, P. canescens, P. cordata, P. coronopus, P. elongata, P. erecta, P. eriopoda, P. firma, P. floccosa, P. helleri, P. heterophylla, P. hookeriana, P. indica, P. lanceolata, P. macrocarpa, P. major, P. maritima, P. media, P. ovata, P. patagonica, P. pusilla, P. rhodosperma, P. rugelii, P. sempervirens, P. sparsiflora, P. subnuda, P. tweedyi, P. virginica, P. wrightiana
Synonyms P. arenaria, P. psyllium P. australis subsp. hirtella, P. hirtella, P. hirtella var. galeottiana, P. hirtella var. mollior
Name authority Linnaeus: Syst. Nat., ed. 10, 2: 896. 1759, legitimacy of — name in question Lamarck: in J. Lamarck and J. L. M. Poiret, Tabl. Encycl. 1: 339. (1792)
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