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plantain des sables, sand plantain

bottlebrush indianwheat, bract plantain, bristly plantain, large bract plantain

Habit Annuals; roots taproots, slender. Annuals; roots taproots, slender.
Stems

100–350 mm, freely branched.

20–40 mm.

Leaves

cauline, opposite, 60–80 × 1–3 mm;

blade linear to linear-lanceolate, margins entire or toothed, veins conspicuous or not, surfaces hairy.

30–200 × 3–7 mm;

blade linear or narrowly lanceolate, margins entire, rarely toothed, veins conspicuous or not, abaxial surface villous, adaxial glabrous or sparsely villous.

Scapes

100–650 mm, hairy.

100–500 mm, hairy, hairs antrorse, long and short.

Spikes

greenish or brownish, (50–)150–250 mm, densely flowered, eglandular;

bracts proximal strongly differing from distal, ovate, 2–5(–7) mm, length 1–1.5 times sepals, proximal bracts: apex acute.

greenish or brownish, 80–150 mm, densely flowered, flowers in whorls or pairs;

bracts almost linear, 15–30 mm, length 2–12 times sepals.

Flowers

sepals 2–3 mm;

corolla radially symmetric, lobes reflexed, 2–4 mm, base obtuse;

stamens 4.

sepals 2.7–3.7 mm;

corolla bilaterally symmetric, lobes reflexed, adaxials 1.4–2.3 mm, laterals symmetric, base deeply cordate;

stamens 4, connective elongated, apex acute.

Seeds

2, 2–2.5 mm.

2, 2.5–2.9 mm.

2n

= 12.

= 20.

Plantago indica

Plantago aristata

Phenology Flowering late summer–fall. Flowering spring–fall.
Habitat Roadsides, railroads, sandy shorelines. Roadsides, pastures, disturbed ground.
Elevation 0–200 m. (0–700 ft.) 0–700 m. (0–2300 ft.)
Distribution
from FNA
CA; CT; DE; IA; IL; IN; MA; ME; MI; MN; MO; NC; NH; NJ; NY; OH; OR; PA; VA; VT; WA; WI; BC; MB; ON; QC; Eurasia [Introduced in North America]
from FNA
AL; AR; CA; CT; DC; DE; FL; GA; IA; IL; IN; KS; KY; LA; MA; MD; ME; MI; MN; MO; MS; NC; NE; NH; NJ; NM; NY; OH; OK; OR; PA; RI; SC; SD; TN; TX; VA; VT; WA; WI; WV; NS; ON [Introduced in Central America, Europe, Asia]
[WildflowerSearch map]
[BONAP county map]
Discussion

While considering the proposal made by W. L. Applequist (2006) to reject Plantago psyllium, the Nomenclature Committee for Vascular Plants also decided that P. indica is a legitimate name (R. K. Brummitt 2009). A. B. Doweld and A. Shipunov (2017) published a proposal to reject P. indica in favor of P. arenaria. That proposal awaits a decision by that committee.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Plantago aristata is similar to P. patagonica; the latter is distinguished by its dense, villous indument.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 17, p. 288. FNA vol. 17, p. 284.
Parent taxa Plantaginaceae > Plantago Plantaginaceae > Plantago
Sibling taxa
P. afra, P. argyrea, P. aristata, P. australis, P. canescens, P. cordata, P. coronopus, P. elongata, P. erecta, P. eriopoda, P. firma, P. floccosa, P. helleri, P. heterophylla, P. hookeriana, P. lanceolata, P. macrocarpa, P. major, P. maritima, P. media, P. ovata, P. patagonica, P. pusilla, P. rhodosperma, P. rugelii, P. sempervirens, P. sparsiflora, P. subnuda, P. tweedyi, P. virginica, P. wrightiana
P. afra, P. argyrea, P. australis, P. canescens, P. cordata, P. coronopus, P. elongata, P. erecta, P. eriopoda, P. firma, P. floccosa, P. helleri, P. heterophylla, P. hookeriana, P. indica, P. lanceolata, P. macrocarpa, P. major, P. maritima, P. media, P. ovata, P. patagonica, P. pusilla, P. rhodosperma, P. rugelii, P. sempervirens, P. sparsiflora, P. subnuda, P. tweedyi, P. virginica, P. wrightiana
Synonyms P. arenaria, P. psyllium P. patagonica var. aristata, P. purshii var. aristata
Name authority Linnaeus: Syst. Nat., ed. 10, 2: 896. 1759, legitimacy of — name in question Michaux: Fl. Bor.-Amer. 1: 95. (1803)
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