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California plantain, Hooker's plantain, tallow weed

blond plantain, desert Indian-wheat, desert plantain

Habit Annuals; roots taproots, slender. Annuals; roots taproots, slender.
Stems

0–10 mm.

0–30 mm, often branched.

Leaves

60–120 × 4–10 mm;

blade linear, margins toothed (teeth to 4 mm), rarely entire, veins conspicuous or not, surfaces villous or sericeous.

10–230 × 0.5–12 mm;

blade linear or narrowly elliptic, margins toothed, veins conspicuous or not, surfaces villous or lanate to sericeous.

Scapes

280–580 mm, hairy, hairs antrorse, long and short.

10–400 mm, hairy, hairs woolly, long.

Spikes

greenish or brownish, 50–180 mm, densely flowered, flowers in whorls or pairs;

bracts ovate or triangular, 1.5–6 mm, length 0.6–1.4 times sepals, apex acute or acuminate.

grayish or brownish, 20–400 mm, densely flowered, flowers in spirals;

bracts ovate or elliptic, 1.7–4 mm, length 0.8–1.2 times sepals, apex not reached by green nerve.

Flowers

sepals 2.5–4 mm;

corolla bilaterally symmetric, abaxial and lateral lobes reflexed, adaxial erect, 2.2–2.5 mm, base slightly cordate;

stamens 4, connective significantly elongated, apex acute.

sepals 1.9–3.5 mm;

corolla radially symmetric, lobes reflexed, 1.3–2.8 mm, base cuneate;

stamens 4.

Seeds

2, 2.4–3.4 mm.

2, 2–2.6 mm.

2n

= 20.

= 8.

Plantago hookeriana

Plantago ovata

Phenology Flowering spring. Flowering spring.
Habitat Sandy soils, disturbed areas. Sandy deserts and steppes.
Elevation 0–1300 m. (0–4300 ft.) 0–1800 m. (0–5900 ft.)
Distribution
from FNA
LA; MS; TX; Mexico (Chihuahua, Coahuila, Nuevo León, Tamaulipas)
[WildflowerSearch map]
[BONAP county map]
from FNA
AZ; CA; NV; TX; UT; Mexico (Baja California, Sonora); Eurasia; Africa
[WildflowerSearch map]
[BONAP county map]
Discussion

From molecular evidence, S. C. Meyers and A. Liston (2008) suggested that Plantago ovata was introduced to North America during the Pleistocene. They recognized four varieties; North American specimens can be treated as two varieties based on bract and corolla color: the inland var. fastigiata (E. Morris) S. C. Meyers & Liston (midribs of mature flower bracts green, corolla lobes without reddish brown midribs) and the coastal var. insularis (Eastwood) S. C. Meyers & Liston (midribs of mature flower bracts brown, corolla lobe midribs prominent, reddish brown). Unfortunately, these features are not easily seen on many herbarium specimens, and these taxa are not recognized here.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 17, p. 287. FNA vol. 17, p. 290.
Parent taxa Plantaginaceae > Plantago Plantaginaceae > Plantago
Sibling taxa
P. afra, P. argyrea, P. aristata, P. australis, P. canescens, P. cordata, P. coronopus, P. elongata, P. erecta, P. eriopoda, P. firma, P. floccosa, P. helleri, P. heterophylla, P. indica, P. lanceolata, P. macrocarpa, P. major, P. maritima, P. media, P. ovata, P. patagonica, P. pusilla, P. rhodosperma, P. rugelii, P. sempervirens, P. sparsiflora, P. subnuda, P. tweedyi, P. virginica, P. wrightiana
P. afra, P. argyrea, P. aristata, P. australis, P. canescens, P. cordata, P. coronopus, P. elongata, P. erecta, P. eriopoda, P. firma, P. floccosa, P. helleri, P. heterophylla, P. hookeriana, P. indica, P. lanceolata, P. macrocarpa, P. major, P. maritima, P. media, P. patagonica, P. pusilla, P. rhodosperma, P. rugelii, P. sempervirens, P. sparsiflora, P. subnuda, P. tweedyi, P. virginica, P. wrightiana
Synonyms P. hookeriana var. nuda P. insularis, P. insularis var. fastigiata
Name authority Fischer & C. A. Meyer: Index Seminum (St. Petersburg) 1838: 39. (1839) Forsskål: Fl. Aegypt.-Arab., 31. (1775)
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