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Heller's plantain

blond plantain, desert Indian-wheat, desert plantain

Habit Annuals; roots taproots, slender. Annuals; roots taproots, slender.
Stems

0–12 mm.

0–30 mm, often branched.

Leaves

50–130 × 3–7 mm;

blade linear, margins entire, veins conspicuous or not, surfaces villous, adaxial rarely glabrous.

10–230 × 0.5–12 mm;

blade linear or narrowly elliptic, margins toothed, veins conspicuous or not, surfaces villous or lanate to sericeous.

Scapes

80–250 mm, hairy, hairs patent, long and antrorse, short.

10–400 mm, hairy, hairs woolly, long.

Spikes

greenish, whitish, or brownish, 50–120 mm, densely flowered, flowers in whorls or pairs;

bracts triangular, 2.5–8 mm, length 0.6–2.2 times sepals.

grayish or brownish, 20–400 mm, densely flowered, flowers in spirals;

bracts ovate or elliptic, 1.7–4 mm, length 0.8–1.2 times sepals, apex not reached by green nerve.

Flowers

sepals 3.5–4 mm;

corolla radially symmetric, lobes reflexed, 3–3.6 mm, base slightly cordate;

stamens 4, connective elongated, apex acute.

sepals 1.9–3.5 mm;

corolla radially symmetric, lobes reflexed, 1.3–2.8 mm, base cuneate;

stamens 4.

Seeds

2, 3.2–3.8 mm.

2, 2–2.6 mm.

2n

= 20.

= 8.

Plantago helleri

Plantago ovata

Phenology Flowering spring. Flowering spring.
Habitat Dry slopes and flats on limestone. Sandy deserts and steppes.
Elevation 100–1700 m. (300–5600 ft.) 0–1800 m. (0–5900 ft.)
Distribution
from FNA
NM; TX
[WildflowerSearch map]
[BONAP county map]
from FNA
AZ; CA; NV; TX; UT; Mexico (Baja California, Sonora); Eurasia; Africa
[WildflowerSearch map]
[BONAP county map]
Discussion

Populations of Plantago helleri are concentrated in central Texas, especially in the Edwards Plateau and Trans-Pecos. The species also occurs in Eddy and Lincoln counties, New Mexico.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

From molecular evidence, S. C. Meyers and A. Liston (2008) suggested that Plantago ovata was introduced to North America during the Pleistocene. They recognized four varieties; North American specimens can be treated as two varieties based on bract and corolla color: the inland var. fastigiata (E. Morris) S. C. Meyers & Liston (midribs of mature flower bracts green, corolla lobes without reddish brown midribs) and the coastal var. insularis (Eastwood) S. C. Meyers & Liston (midribs of mature flower bracts brown, corolla lobe midribs prominent, reddish brown). Unfortunately, these features are not easily seen on many herbarium specimens, and these taxa are not recognized here.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 17, p. 287. FNA vol. 17, p. 290.
Parent taxa Plantaginaceae > Plantago Plantaginaceae > Plantago
Sibling taxa
P. afra, P. argyrea, P. aristata, P. australis, P. canescens, P. cordata, P. coronopus, P. elongata, P. erecta, P. eriopoda, P. firma, P. floccosa, P. heterophylla, P. hookeriana, P. indica, P. lanceolata, P. macrocarpa, P. major, P. maritima, P. media, P. ovata, P. patagonica, P. pusilla, P. rhodosperma, P. rugelii, P. sempervirens, P. sparsiflora, P. subnuda, P. tweedyi, P. virginica, P. wrightiana
P. afra, P. argyrea, P. aristata, P. australis, P. canescens, P. cordata, P. coronopus, P. elongata, P. erecta, P. eriopoda, P. firma, P. floccosa, P. helleri, P. heterophylla, P. hookeriana, P. indica, P. lanceolata, P. macrocarpa, P. major, P. maritima, P. media, P. patagonica, P. pusilla, P. rhodosperma, P. rugelii, P. sempervirens, P. sparsiflora, P. subnuda, P. tweedyi, P. virginica, P. wrightiana
Synonyms P. insularis, P. insularis var. fastigiata
Name authority Small: Bull. New York Bot. Gard. 1: 288. (1899) Forsskål: Fl. Aegypt.-Arab., 31. (1775)
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