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Heller's plantain

hoary plantain, plantain moyen

Habit Annuals; roots taproots, slender. Perennials; caudex glabrous or hairy; roots taproots, thick.
Stems

0–12 mm.

0–20 mm.

Leaves

50–130 × 3–7 mm;

blade linear, margins entire, veins conspicuous or not, surfaces villous, adaxial rarely glabrous.

prostrate, sometimes ascending, 40–200 × 30–70 mm;

blade elliptic to ovate, margins entire or toothed, veins conspicuous, surfaces hairy, hairs 0.5 mm.

Scapes

80–250 mm, hairy, hairs patent, long and antrorse, short.

80–300 mm, not groove-angled, hirsute.

Spikes

greenish, whitish, or brownish, 50–120 mm, densely flowered, flowers in whorls or pairs;

bracts triangular, 2.5–8 mm, length 0.6–2.2 times sepals.

grayish or whitish, 100–400 mm, densely flowered, shiny;

corolla lobes of neighboring flowers overlapping;

bracts ovate, 1.8–2 mm, length 0.9–1 times sepals.

Flowers

sepals 3.5–4 mm;

corolla radially symmetric, lobes reflexed, 3–3.6 mm, base slightly cordate;

stamens 4, connective elongated, apex acute.

sepals 2 mm, adaxial 2 nearly distinct;

corolla radially symmetric, lobes reflexed, 1.5 mm, base obtuse;

stamens 4.

Seeds

2, 3.2–3.8 mm.

2–4, 2 mm.

2n

= 20.

= 12, 24.

Plantago helleri

Plantago media

Phenology Flowering spring. Flowering summer.
Habitat Dry slopes and flats on limestone. Disturbed areas, neutral and basic soils.
Elevation 100–1700 m. (300–5600 ft.) 0–200 m. (0–700 ft.)
Distribution
from FNA
NM; TX
[WildflowerSearch map]
[BONAP county map]
from FNA
CT; IL; MA; ME; MI; NH; NJ; NY; PA; RI; WI; MB; NB; ON; QC; Eurasia [Introduced in North America]
[BONAP county map]
Discussion

Populations of Plantago helleri are concentrated in central Texas, especially in the Edwards Plateau and Trans-Pecos. The species also occurs in Eddy and Lincoln counties, New Mexico.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Three varieties of Plantago media have been recognized in western Europe. It is not clear how names of those varieties apply to P. media in eastern Europe and North America.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 17, p. 287. FNA vol. 17, p. 290.
Parent taxa Plantaginaceae > Plantago Plantaginaceae > Plantago
Sibling taxa
P. afra, P. argyrea, P. aristata, P. australis, P. canescens, P. cordata, P. coronopus, P. elongata, P. erecta, P. eriopoda, P. firma, P. floccosa, P. heterophylla, P. hookeriana, P. indica, P. lanceolata, P. macrocarpa, P. major, P. maritima, P. media, P. ovata, P. patagonica, P. pusilla, P. rhodosperma, P. rugelii, P. sempervirens, P. sparsiflora, P. subnuda, P. tweedyi, P. virginica, P. wrightiana
P. afra, P. argyrea, P. aristata, P. australis, P. canescens, P. cordata, P. coronopus, P. elongata, P. erecta, P. eriopoda, P. firma, P. floccosa, P. helleri, P. heterophylla, P. hookeriana, P. indica, P. lanceolata, P. macrocarpa, P. major, P. maritima, P. ovata, P. patagonica, P. pusilla, P. rhodosperma, P. rugelii, P. sempervirens, P. sparsiflora, P. subnuda, P. tweedyi, P. virginica, P. wrightiana
Name authority Small: Bull. New York Bot. Gard. 1: 288. (1899) Linnaeus: Sp. Pl. 1: 113. (1753)
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