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Heller's plantain

Alaska plantain, seashore plantain

Habit Annuals; roots taproots, slender. Perennials; caudex well developed, conspicuous, glabrous; roots fibrous, thick.
Stems

0–12 mm.

0–20 mm.

Leaves

50–130 × 3–7 mm;

blade linear, margins entire, veins conspicuous or not, surfaces villous, adaxial rarely glabrous.

(80–)100–400(–550) × (5–)10–35(–40) mm;

blade oblanceolate or almost linear, margins entire, veins conspicuous, 4 or 5, surfaces glabrous.

Scapes

80–250 mm, hairy, hairs patent, long and antrorse, short.

300–400 mm, glabrous or sparsely hairy, becoming densely so distally.

Spikes

greenish, whitish, or brownish, 50–120 mm, densely flowered, flowers in whorls or pairs;

bracts triangular, 2.5–8 mm, length 0.6–2.2 times sepals.

greenish or brownish, 350–450 mm, loosely flowered;

bracts ovate to deltate, 3–4 mm, length 1.5–2.5 times sepals.

Flowers

sepals 3.5–4 mm;

corolla radially symmetric, lobes reflexed, 3–3.6 mm, base slightly cordate;

stamens 4, connective elongated, apex acute.

sepals 1.5–2 mm;

corolla radially symmetric, lobes spreading, 1.5–2 mm, base obtuse;

stamens 4.

Fruits

ovoid, indehiscent or dehiscence not circumscissile.

Seeds

2, 3.2–3.8 mm.

1 or 2, 4–5 mm.

2n

= 20.

= 24.

Plantago helleri

Plantago macrocarpa

Phenology Flowering spring. Flowering late spring–early summer.
Habitat Dry slopes and flats on limestone. Wet places, tidal marshes, saline areas.
Elevation 100–1700 m. (300–5600 ft.) 0–700 m. (0–2300 ft.)
Distribution
from FNA
NM; TX
[WildflowerSearch map]
[BONAP county map]
from FNA
AK; OR; WA; BC; Asia
[WildflowerSearch map]
[BONAP county map]
Discussion

Populations of Plantago helleri are concentrated in central Texas, especially in the Edwards Plateau and Trans-Pecos. The species also occurs in Eddy and Lincoln counties, New Mexico.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Plantago macrocarpa has been documented along the Pacific coast south to the mouth of the Yachats River in Oregon.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 17, p. 287. FNA vol. 17, p. 289.
Parent taxa Plantaginaceae > Plantago Plantaginaceae > Plantago
Sibling taxa
P. afra, P. argyrea, P. aristata, P. australis, P. canescens, P. cordata, P. coronopus, P. elongata, P. erecta, P. eriopoda, P. firma, P. floccosa, P. heterophylla, P. hookeriana, P. indica, P. lanceolata, P. macrocarpa, P. major, P. maritima, P. media, P. ovata, P. patagonica, P. pusilla, P. rhodosperma, P. rugelii, P. sempervirens, P. sparsiflora, P. subnuda, P. tweedyi, P. virginica, P. wrightiana
P. afra, P. argyrea, P. aristata, P. australis, P. canescens, P. cordata, P. coronopus, P. elongata, P. erecta, P. eriopoda, P. firma, P. floccosa, P. helleri, P. heterophylla, P. hookeriana, P. indica, P. lanceolata, P. major, P. maritima, P. media, P. ovata, P. patagonica, P. pusilla, P. rhodosperma, P. rugelii, P. sempervirens, P. sparsiflora, P. subnuda, P. tweedyi, P. virginica, P. wrightiana
Name authority Small: Bull. New York Bot. Gard. 1: 288. (1899) Chamisso & Schlechtendal: Linnaea 1: 166. (1826)
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