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Heller's plantain

plantain des sables, sand plantain

Habit Annuals; roots taproots, slender. Annuals; roots taproots, slender.
Stems

0–12 mm.

100–350 mm, freely branched.

Leaves

50–130 × 3–7 mm;

blade linear, margins entire, veins conspicuous or not, surfaces villous, adaxial rarely glabrous.

cauline, opposite, 60–80 × 1–3 mm;

blade linear to linear-lanceolate, margins entire or toothed, veins conspicuous or not, surfaces hairy.

Scapes

80–250 mm, hairy, hairs patent, long and antrorse, short.

100–650 mm, hairy.

Spikes

greenish, whitish, or brownish, 50–120 mm, densely flowered, flowers in whorls or pairs;

bracts triangular, 2.5–8 mm, length 0.6–2.2 times sepals.

greenish or brownish, (50–)150–250 mm, densely flowered, eglandular;

bracts proximal strongly differing from distal, ovate, 2–5(–7) mm, length 1–1.5 times sepals, proximal bracts: apex acute.

Flowers

sepals 3.5–4 mm;

corolla radially symmetric, lobes reflexed, 3–3.6 mm, base slightly cordate;

stamens 4, connective elongated, apex acute.

sepals 2–3 mm;

corolla radially symmetric, lobes reflexed, 2–4 mm, base obtuse;

stamens 4.

Seeds

2, 3.2–3.8 mm.

2, 2–2.5 mm.

2n

= 20.

= 12.

Plantago helleri

Plantago indica

Phenology Flowering spring. Flowering late summer–fall.
Habitat Dry slopes and flats on limestone. Roadsides, railroads, sandy shorelines.
Elevation 100–1700 m. (300–5600 ft.) 0–200 m. (0–700 ft.)
Distribution
from FNA
NM; TX
[WildflowerSearch map]
[BONAP county map]
from FNA
CA; CT; DE; IA; IL; IN; MA; ME; MI; MN; MO; NC; NH; NJ; NY; OH; OR; PA; VA; VT; WA; WI; BC; MB; ON; QC; Eurasia [Introduced in North America]
Discussion

Populations of Plantago helleri are concentrated in central Texas, especially in the Edwards Plateau and Trans-Pecos. The species also occurs in Eddy and Lincoln counties, New Mexico.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

While considering the proposal made by W. L. Applequist (2006) to reject Plantago psyllium, the Nomenclature Committee for Vascular Plants also decided that P. indica is a legitimate name (R. K. Brummitt 2009). A. B. Doweld and A. Shipunov (2017) published a proposal to reject P. indica in favor of P. arenaria. That proposal awaits a decision by that committee.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 17, p. 287. FNA vol. 17, p. 288.
Parent taxa Plantaginaceae > Plantago Plantaginaceae > Plantago
Sibling taxa
P. afra, P. argyrea, P. aristata, P. australis, P. canescens, P. cordata, P. coronopus, P. elongata, P. erecta, P. eriopoda, P. firma, P. floccosa, P. heterophylla, P. hookeriana, P. indica, P. lanceolata, P. macrocarpa, P. major, P. maritima, P. media, P. ovata, P. patagonica, P. pusilla, P. rhodosperma, P. rugelii, P. sempervirens, P. sparsiflora, P. subnuda, P. tweedyi, P. virginica, P. wrightiana
P. afra, P. argyrea, P. aristata, P. australis, P. canescens, P. cordata, P. coronopus, P. elongata, P. erecta, P. eriopoda, P. firma, P. floccosa, P. helleri, P. heterophylla, P. hookeriana, P. lanceolata, P. macrocarpa, P. major, P. maritima, P. media, P. ovata, P. patagonica, P. pusilla, P. rhodosperma, P. rugelii, P. sempervirens, P. sparsiflora, P. subnuda, P. tweedyi, P. virginica, P. wrightiana
Synonyms P. arenaria, P. psyllium
Name authority Small: Bull. New York Bot. Gard. 1: 288. (1899) Linnaeus: Syst. Nat., ed. 10, 2: 896. 1759, legitimacy of — name in question
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