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glandular plantain

common plantain, great plantain, nippleseed, plantain majeur

Habit Annuals; roots taproots, slender. Perennials, sometimes annuals; caudex absent; roots fibrous, thick.
Stems

100–350 mm, freely branched.

0–20 mm.

Leaves

cauline, opposite, 30–60 × 1–4 mm;

blade linear to linear-lanceolate, margins entire or slightly toothed, veins conspicuous or not, surfaces hairy.

20–150(–400) × 10–120(–170) mm;

petiole to 200 mm;

blade ovate to cordate-ovate, margins entire or toothed, veins conspicuous, surfaces glabrous or hirsute.

Scapes

30–50 mm, hairy.

50–250(–500) mm, surpassing leaves or not, glabrous or hirsute.

Spikes

greenish or brownish, 40–65 mm, densely flowered, glandular-hairy;

bracts all similar, ovate, 3–5 mm, lengths 1–1.5 times sepals.

brownish or greenish, (20–)50–300(–400) mm, densely flowered;

bracts lanceolate, 0.5–1 mm, length 0.3–0.7 times sepals.

Flowers

sepals 3–3.5 mm;

corolla radially symmetric, lobes reflexed, 2–3 mm, base obtuse;

stamens 4.

sepals 1.5–2 mm;

corolla radially symmetric, lobes reflexed, 0.5–1 mm, base obtuse;

stamens 4.

Fruits

(2–)4–5 mm, dehiscing at middle.

Capsules

lanceoloid.

Seeds

2, 2–3 mm.

5–35, 0.5–1 mm.

2n

= 12.

= 12.

Plantago afra

Plantago major

Phenology Flowering summer. Flowering summer.
Habitat Disturbed habitats. Roadsides, trails, stream banks, urban areas, lawns, other disturbed areas.
Elevation 0–200 m. (0–700 ft.) 0–3000 m. (0–9800 ft.)
Distribution
from FNA
MA; s Europe [Introduced in North America]
from FNA
AK; AL; AR; AZ; CA; CO; CT; DC; DE; FL; GA; IA; ID; IL; IN; KS; KY; LA; MA; MD; ME; MI; MN; MO; MS; MT; NC; ND; NE; NH; NJ; NM; NV; NY; OH; OK; OR; PA; RI; SC; SD; TN; TX; UT; VA; VT; WA; WI; WV; WY; AB; BC; MB; NB; NL; NS; NT; ON; PE; QC; SK; YT; SPM; Greenland; Eurasia [Introduced in North America; introduced also in Mexico, Central America, South America, Africa, Pacific Islands, Australia]
[WildflowerSearch map]
[BONAP county map]
Discussion

Plantago afra is known in Massachusetts from a single collection made in 1927 in Worcester County.

Plantago psyllium Linnaeus (1762, not 1753), a rejected name, and P. indica Linnaeus are misapplied names that pertain here. Plantago squalida Salisbury is an illegitimate name that pertains here.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Subspecies intermedia (Gilibert) Lange is often accepted by European botanists as a separate species, Plantago uliginosa F. W. Schmidt. Observations suggest that plants referable to this taxon may occur in the United States; it has been reported from the New England states (A. Haines 2011). Subspecies intermedia is distinguished by more abundant (11–35) and smaller (0.8–1 mm) seeds, ascending spikes, ovoid fruits, and elliptic or lanceolate leaf blades. Without detailed morphologic and genetic investigations of North American plants similar to the study of European plants by M. Morgan-Richards and K. Wolff (1999), it is not possible to draw any conclusions about the status and distribution of this or any other possible infraspecific taxa of P. major in North America.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 17, p. 284. FNA vol. 17, p. 289.
Parent taxa Plantaginaceae > Plantago Plantaginaceae > Plantago
Sibling taxa
P. argyrea, P. aristata, P. australis, P. canescens, P. cordata, P. coronopus, P. elongata, P. erecta, P. eriopoda, P. firma, P. floccosa, P. helleri, P. heterophylla, P. hookeriana, P. indica, P. lanceolata, P. macrocarpa, P. major, P. maritima, P. media, P. ovata, P. patagonica, P. pusilla, P. rhodosperma, P. rugelii, P. sempervirens, P. sparsiflora, P. subnuda, P. tweedyi, P. virginica, P. wrightiana
P. afra, P. argyrea, P. aristata, P. australis, P. canescens, P. cordata, P. coronopus, P. elongata, P. erecta, P. eriopoda, P. firma, P. floccosa, P. helleri, P. heterophylla, P. hookeriana, P. indica, P. lanceolata, P. macrocarpa, P. maritima, P. media, P. ovata, P. patagonica, P. pusilla, P. rhodosperma, P. rugelii, P. sempervirens, P. sparsiflora, P. subnuda, P. tweedyi, P. virginica, P. wrightiana
Synonyms P. major var. pachyphylla, P. major var. pilgeri, P. major var. scopulorum
Name authority Linnaeus: Sp. Pl. ed. 2, 1: 168. (1762) Linnaeus: Sp. Pl. 1: 112. (1753)
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