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pinon ricegrass, pinyon ricegrass

speargrass

Habit Plants perennial; cespitose, not rhizomatous.
Culms

35-95 cm, usually glabrous, sometimes pubescent below the nodes;

nodes 2-3, often dark, glabrous.

4-150 cm, usually erect, sometimes decumbent, glabrous, not branched above the base;

basal branching intravaginal;

prophylls shorter than the sheaths, mostly glabrous, keels usually with hairs, apices bifid, teeth 1-3 mm;

cleistogenes not developed.

Sheaths

glabrous, smooth;

ligules truncate to rounded, of basal leaves 0.4-1.8 mm, of upper leaves 1.5-2 mm;

blades 6-26 cm long, usually involute and 0.3-5 mm in diameter, sometimes flat and 0.5-1(1.5) mm wide, 3-veined, both surfaces glabrous, veins often scabridulous, margins scabrous.

open to the base, margins glabrous;

ligules membranous, decurrent, truncate to acute, sometimes highest at the sides, sometimes ciliate;

blades convolute to flat, translucent between the veins, often sinuous distally.

Panicles

6.5-25 cm, open, often partially enclosed in the upper leaf sheath, with 20-60 spikelets;

branches flexuous;

pedicels 4-12 mm, flattened, hispid.

Inflorescences

terminal panicles, open or contracted, spikelets usually confined to the distal 1/2 of each branch.

Spikelets

4-22 mm, with 1 floret;

rachillas not prolonged beyond the base of the floret;

disarticulation above the glumes, beneath the floret.

Glumes

subequal, 4-6.2 mm long, 1.8-3.1 mm wide, 5-7-veined, often partly purplish;

florets 3-5.5 mm long, 0.6-1.9 mm thick, somewhat laterally compressed, rectangular to slightly obovate in side view;

calluses 0.2-0.7 mm, blunt, strigose;

lemmas tan to light chocolate brown, shiny, smooth, evenly pubescent when immature, hairs easily rubbed off;

crowns about 0.8 mm wide, inconspicuous, glabrous or glabrate;

awns 11-20 mm, persistent, twice-geniculate;

paleas about 3.5 mm;

lodicules 2, about 1 mm;

anthers 0.3-0.5 mm, not penicillate.

subequal, longer than the floret, lanceolate, 3-7(8)-veined;

florets globose to fusiform, terete to laterally compressed;

calluses well developed, sharp or blunt, glabrous or antrorsely strigose, hairs yellow to golden brown;

lemmas coriaceous to indurate, glabrous or pubescent, striate, particularly near the base, smooth, papillose, or tuberculate, often smooth on the lower portion and papillate to tuberculate distally, margins involute, fitting into the grooved palea, apices fused into a crown, awned, lemmas often narrowed below the crown, crowns usually ciliate;

awns caducous to persistent, usually twice-geniculate, first 2 segments usually twisted and hispid, terminal segment straight and scabridulous;

paleas longer than the lemmas, similar in texture, glabrous, sulcate between the veins, apices prow-tipped;

lodicules 2 or 3, membranous, glabrous, blunt or acute;

anthers 3;

ovaries glabrous;

styles 2.

Caryopses

2.5-3 mm long, about 0.6 mm thick, fusiform.

terete to globose or lens-shaped, x = 11.

2n

= 42.

Piptochaetium fimbriatum

Piptochaetium

Distribution
from FNA
AZ; NM; TX
[WildflowerSearch map]
[BONAP county map]
from FNA
AL; AR; AZ; CA; CT; DC; DE; FL; GA; IN; KY; LA; MA; MD; MI; MO; MS; NC; NJ; NM; NY; OH; OK; PA; RI; SC; TN; TX; VA; WV; ON
[BONAP county map]
Discussion

Piptochaetium fimbriatum is an attractive species that grows in oak and pinyon woods of the southwestern United States and adjacent Mexico, and merits consideration as an ornamental. It has also been reported from Guatemala; the report has not been verified.

Piptochaetium fimbriatum is not easily confused with other species in our range. Hitchcock (1951) treated it as including P. seleri (Pilg.) Henrard, a Mexican species with dull, rough, oblanceolate lemmas and persistent lemma hairs, an interpretation that is no longer accepted. It is occasionally confused with P. pringlei; it differs from that species in having shorter florets and blunt calluses.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Piptochaetium is primarily South American, being particularly abundant in Argentina. It has 27 species. Four species are native in the Flora region; two South American species are established at a single location in Marin County, California.

All stipoid species with elongate florets were included in Stipa by Hitchcock (1935, 1951) and other North American taxonomists. Parodi (1944) argued that Piptochaetium should be expanded to include those species of Stipa with elongated florets that shared with Piptochaetium its distinctive lemma and palea morphology, hair color, and leaf anatomy. His interpretation is now universally accepted.

The basal chromosome number of Piptochaetium is probably 11. This interpretation implies that counts of In = 42 (Gould 1965 and Reeder 1968 for P. fimbriatum; Reeder 1977 for P. pringlei) represent an aneuploid reduction from 2n = 44. It is also possible that the base number is 7, as is common in the Pooideae.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Key
1. Florets 6.5-22 mm long; culms 40-130 cm tall.
→ 2
2. Lemmas hairy; awns 19-35 mm long
P. pringlei
2. Lemmas glabrous; awns 40-120 mm long.
→ 3
3. Florets 7-13 mm long; awns 40-75 mm long
P. avenaceum
3. Florets 13.5-22 mm long; awns 62-120 mm long
P. avenacioides
1. Florets 2.3-5.5 mm long; culms 20-95 cm tall.
→ 4
4. Lemmas golden brown, hairy, the hairs easily rubbed off
P. fimbriatum
4. Lemmas dark brown, glabrous.
→ 5
5. Awns 10-16 mm long; blades 0.8-1.5 mm wide; distal margin of the lemma crowns straight
P. setosum
5. Awns 15-25 mm long; blades 0.2-0.4 mm wide; distal margin of the lemma crowns sometimes slightly to strongly revolute
P. stipoides
Source FNA vol. 24. FNA vol. 24, p. 161. Author: Mary E. Barkworth;.
Parent taxa Poaceae > subfam. Pooideae > tribe Stipeae > Piptochaetium Poaceae > subfam. Pooideae > tribe Stipeae
Sibling taxa
P. avenaceum, P. avenacioides, P. pringlei, P. setosum, P. stipoides
Subordinate taxa
P. avenaceum, P. avenacioides, P. fimbriatum, P. pringlei, P. setosum, P. stipoides
Name authority (Kunth) Hitchc. J. Presl
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