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smilo grass

Shoshone piptatherum

Habit Plants loosely cespitose, not rhizomatous. Plants tightly cespitose, not rhizomatous.
Culms

40-150 cm, glabrous, often branching at the lower cauline nodes;

basal branching extravaginal.

20-50 cm, internodes smooth to scabridulous;

nodes glabrate;

basal branching intravaginal.

Leaves

not basally concentrated;

sheaths glabrous, persistent;

ligules of lower leaves 0.5-1.5 mm, truncate;

ligules of upper leaves 1.5-4 mm, rounded to sharply acute;

blades 5-30 cm long, 2-10 mm wide, flat, smooth on both surfaces.

basally concentrated;

ligules 1.8-5.5 mm, hyaline, acute, often lacerate;

blades 4-16 cm long, 1-2.5 mm wide when flat, 0.6-1 mm in diameter when involute, abaxial surfaces smooth or scabridulous, adaxial surfaces scabridulous.

Panicles

10-40 cm, lax, lower nodes either with 3-7 branches bearing 10-40 functional spikelets, or with 15-30+ branches with no functional spikelets;

primary branches spreading to ascending;

lower branches 3-8 cm;

secondary branches diverging from the primary branches.

3.3-22 cm, lower nodes with 1-2(4) branches;

branches 1.8-11.6 cm, flexuous, initially appressed, becoming strongly divergent to reflexed, secondary and tertiary branches appressed to the primary branches.

Glumes

2.5-3.5 mm, acuminate, 3-veined;

florets 1.5-2 mm, dorsally compressed;

calluses about 0.3 mm, glabrous, disarticulation scars circular;

lemmas stiffly membranous, glabrous, margins fused at the base, not overlapping, light brown at maturity;

awns 3-4 mm;

anthers 2-2.5 mm, penicillate;

ovaries rounded, bearing two styles.

subequal, 3.2-5.3 mm, exceeding the florets by 0.2-1.5 mm, ovate to broadly lanceolate, 1-9-veined, apices acute to acuminate;

florets 2.4-4.1 mm, dorsally compressed;

calluses about 0.3 mm, hairy, disarticulation scars round;

lemmas coriaceous, evenly pubescent, hairs to 0.5 mm, becoming tawny with age, margins not overlapping at maturity;

awns 1-2.5 mm, straight or slightly arcuate, caducous;

paleas 2.1-3.6 mm, similar to the lemmas in texture and pubescence;

anthers 1.7-2.2 mm, penicillate;

ovaries truncate, bearing 2 styles.

Caryopses

1.5-1.7 mm long, about 0.8 mm thick;

hila linear, about 1/2 as long as the caryopses.

1.8-2 mm long, about 0.8 mm thick;

hila linear, about 9/10 the length of the caryopses.

2n

= 24.

= 20.

Piptatherum miliaceum

Piptatherum shoshoneanum

Distribution
from FNA
AZ; CA; ID; MD; NJ; NV; PA; UT
[WildflowerSearch map]
[BONAP county map]
from USDA
Discussion

Piptatherum miliaceum is a Eurasian introduction that is now established in several parts of the world. In its native range it grows, often as a common species, primarily in disturbed areas, wadis, and oases, penetrating into the semidesert regions of northern Africa and western Asia. It is used as a fodder plant in northern Africa. Within the Flora region, P. miliaceum is known from Arizona and California, growing in disturbed sites. It has also been found on a ballast dump in Maryland.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Piptatherum shoshoneanum is known best from eastern Idaho, where it grows in the canyons of the Middle Fork of the Salmon River and its tributaries. It has also been found 750 km to the southwest in the Belted Range of southwestern Nevada. So far, fieldwork on the intervening mountain ranges has not revealed additional populations. It usually grows in moist crevices of igneous, metamorphic, or sedimentary cliffs and rock walls.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Key
1. Panicle branches loosely whorled, lower nodes with 3-7 branches, all spikelet-bearing
subsp. miliaceum
1. Panicle branches densely whorled, lower nodes with 15-30+ branches, some with highly reduced or no spikelets
subsp. thomasii
Source FNA vol. 24, p. 151. FNA vol. 24.
Parent taxa Poaceae > subfam. Pooideae > tribe Stipeae > Piptatherum Poaceae > subfam. Pooideae > tribe Stipeae > Piptatherum
Sibling taxa
P. canadense, P. exiguum, P. micranthum, P. pungens, P. racemosum, P. shoshoneanum
P. canadense, P. exiguum, P. micranthum, P. miliaceum, P. pungens, P. racemosum
Subordinate taxa
P. miliaceum subsp. miliaceum, P. miliaceum subsp. thomasii
Synonyms Oryzopsis miliacea
Name authority (L.) Coss. (Curto & Douglass M. Hend.) P.M. Peterson & Soreng
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