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Canada mountain-rice grass, Canadian piptatherum, Canadian ricegrass, oryzopsis du Canada

ricegrass

Habit Plants cespitose, not rhizomatous. Plants perennial; cespitose or soboliferous, sometimes rhizomatous.
Culms

30-90 cm, glabrous;

basal branching mostly intravaginal.

10-140(150) cm, erect, usually glabrous, usually smooth;

nodes 1-6; branching intra- or extravaginal at the base, not branching above the base;

prophylls concealed by the leaf sheaths.

Leaves

basally concentrated;

sheaths smooth or scabridulous;

ligules 1-4 mm, hyaline, truncate, rounded, or acute;

basal blades 4-15 cm long, 1-1.5 mm wide when flat, 0.5-0.8 mm in diameter when folded or convolute.

sometimes basally concentrated;

cleistogenes not present;

sheaths open, glabrous, smooth to scabrous;

auricles absent;

ligules 0.2-15 mm, membranous to hyaline;

blades 0.5-16 mm wide, flat, involute, valvate, or folded, often tapering in the distal 1/3, apices acute to acuminate, not stiff, basal blades not overwintering, sometimes not developed, flag leaf blades well developed, longer than 1 cm.

Panicles

9-15 cm, lower nodes with 1-2 branches;

branches 1-6 cm, somewhat flexuous, ascending to divergent.

Inflorescences

3-40 cm, terminal panicles, open or contracted;

branches straight or flexuous, usually scabrous, rarely smooth;

pedicels often appressed to the branches.

Spikelets

1.5-7.5 mm, with 1 floret;

rachillas not prolonged beyond the floret;

disarticulation above the glumes, beneath the floret.

Glumes

subequal, 3-6 mm long, 1.3-2 mm wide, ovate, 1-3-veined, apices acute to mucronate;

florets 2.2-4.5 mm, obovoid, dorsally compressed;

calluses 0.2-0.5 mm, hairy, disarticulation scars elliptic;

lemmas coriaceous, evenly pubescent, tan at maturity, margins widely separated even when immature;

awns 5-15 mm, persistent, once- or twice-geniculate, first segments strongly twisted;

paleas similar to the lemmas in length, texture, and pubescence;

anthers 1-2 mm;

ovaries developing 2 conelike style bases, each bearing a single, unbranched style.

from 1 mm shorter than to exceeding the florets, subequal or the lower glumes longer than the upper glumes, membranous, 1-9-veined, veins evident, apices obtuse to acute or acuminate;

florets 1.5-10 mm, usually dorsally compressed, sometimes terete;

calluses 0.1-0.6 mm, glabrous or with hairs, blunt;

lemmas 1.2-9 mm, smooth, coriaceous or stiffly membranous, tawny or light brown to black at maturity, 3-7-veined, margins flat, separated and parallel for their whole length at maturity, apices not lobed or lobed, glabrous or hairy, hairs about 0.5 mm, not spreading, awned, lemma-awn junction evident;

awns 1-18(20) mm, centric, often caducous, almost straight to once- or twice-geniculate, scabrous;

paleas as long as or slightly longer than the lemmas, similar in texture and pubescence, 2(3)-veined, not keeled over the veins, flat between the veins, veins terminating near the apices, apices often pinched;

anthers 3, 0.6-5 mm, sometimes penicillate;

styles 2 and free to their bases, or 1 with 2-3 branches.

Caryopses

about 2.5 mm long, 0.5 mm thick;

hila linear, almost equaling the caryopses.

glabrous, ovoid to obovoid;

hila 1/2 as long as to equaling the length of the caryopses.

x

= 11, 12.

2n

= 22.

Piptatherum canadense

Piptatherum

Distribution
from USDA
[WildflowerSearch map]
from FNA
AZ; CA; ID; MD; NJ; NV; PA; UT
[BONAP county map]
Discussion

Piptatherum canadense grows in grasslands and open woods, from the British Columbia-Alberta border east to Newfoundland, extending south into the Great Lakes region and the northeastern United States. Its persistent, longer awns distinguish P. canadense from P. pungens.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Piptatherum has approximately 30 species, most of which are Eurasian. They extend from lowland to alpine regions, and grow in habitats ranging from mesic forests to semideserts.

The pistils in Piptatherum exhibit variability in the development of the styles, a feature that can be seen only in florets shortly before or at anthesis. This variability is reported in the descriptions, but the number of specimens examined per species is low, sometimes only one.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Key
1. Basal leaf blades 0-2 cm long; cauline leaf blades 8-16 mm wide; florets 4.5-7.5 mm long
P. racemosum
1. Basal leaf blades 4-45 cm long; cauline leaf blades 0.5-10 mm wide; florets 1.5-6 mm long.
→ 2
2. Lemmas and calluses usually glabrous, occasionally sparsely pubescent; florets 1.5-2.5 mm long.
→ 3
3. Blades 0.5-2.5 mm wide, often involute; panicles 5-20 cm long, the lower nodes with 1-3 branches
P. micranthum
3. Blades 2-10 mm wide, flat; panicles 10-40 cm long, the lower nodes usually with 3-7 branches, sometimes with 15-30+ branches
P. miliaceum
2. Lemmas evenly pubescent; calluses hairy; florets 1.5-6 mm long.
→ 4
4. Awns 3.9-15 mm long, persistent, once- or twice-geniculate.
→ 5
5. Primary panicle branches straight, appressed; awns 3.9-7 mm long; florets 3-6 mm long
P. exiguum
5. Primary panicle branches somewhat flexuous, often divergent; awns 5-15 mm long; florets 2.2-4.5 mm long
P. canadense
4. Awns 1-8 mm long, caducous, often absent from herbarium specimens, straight or arcuate.
→ 6
6. Awns 4-8 mm long; florets 1.5-2.5 mm long
P. micranthum
6. Awns 1-2.5 mm long; florets 2.2-4.5 mm long.
→ 7
7. Lower panicle branches straight; ligules 0.5-2.5 mm long
P. pungens
7. Lower panicle branches flexuous; ligules 1.8-5.5 mm long
P. shoshoneanum
Source FNA vol. 24, p. 146. FNA vol. 24, p. 144. Author: Mary E. Barkworth;.
Parent taxa Poaceae > subfam. Pooideae > tribe Stipeae > Piptatherum Poaceae > subfam. Pooideae > tribe Stipeae
Sibling taxa
P. exiguum, P. micranthum, P. miliaceum, P. pungens, P. racemosum, P. shoshoneanum
Subordinate taxa
P. canadense, P. exiguum, P. micranthum, P. miliaceum, P. pungens, P. racemosum, P. shoshoneanum
Synonyms Oryzopsis canadensis
Name authority (Poir.) Dorn P. Beauv.
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