Pinus rigida |
Pinaceae |
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pin rigide, pitch pine |
pine family |
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Habit | Trees to 31m; trunk to 0.9m diam., straight or crooked, commonly with adventitious sprouts; crown rounded or irregular. | Trees (occasionally shrubs), evergreen (annually deciduous in Larix), resinous and aromatic, monoecious. | ||||||||||||||||||||||||
Roots | fibrous to woody, unspecialized. |
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Bark | red-brown, deeply and irregularly furrowed, with long, irregularly rectangular, flat, scaly ridges, resin pockets absent. |
smooth to scaly or furrowed. |
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Branches | arching-spreading to ascending, poorly self-pruning; 2-year-old branchlets stout (mostly over 5mm thick), orange-brown, aging darker brown, rough. |
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Lateral branches | well developed and similar to leading (long) shoots or reduced to well-defined short (spur) shoots (Pinus, Larix); twigs terete, sometimes clothed by persistent primary leaves or leaf bases; longest internodes less than 1cm; buds conspicuous. |
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Buds | ovoid to ovoid-cylindric, red-brown, ca. 1–1.5cm, resinous; scale margins fringed, apex cuspidate. |
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Leaves | 3(–5) per fascicle, spreading to ascending, persisting 2–3 years, 5–10(–15)cm × 1–1.5(–2)mm, straight, twisted, deep to pale yellow-green, all surfaces with fine stomatal lines, margins serrulate, apex abruptly subulate-acuminate; sheath 0.9–1.2cm, base persistent. |
(needles) simple, shed singly (except whole fascicles shed in Pinus), alternate and spirally arranged but sometimes proximally twisted so as to appear 1- or 2-ranked, or fascicled, linear to needlelike, sessile to short-petiolate; foliage leaves either borne singly (spirally) on long shoots or in tufts (fascicles) on short shoots; juvenile leaves (when present) borne on long shoots, scalelike; resin canals present. |
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Pollen cones | cylindric, ca. 20mm, yellow. |
maturing and shed annually, solitary or clustered, axillary, ovoid to ellipsoid or cylindric; sporophylls overlapping, bearing 2 abaxial microsporangia (pollen sacs); pollen spheric, 2-winged, less commonly with wings reduced to frill (in Tsuga sect. Tsuga), or not winged (in Larix and Pseudotsuga). |
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Seed(s) | cones maturing in 2 years, shedding seeds soon thereafter or variously serotinous and long-persistent, often clustered, symmetric, conic to ovoid before opening, broadly ovoid with flat or slightly convex base when open, 3–9cm, creamy brown to light red-brown, sessile to short-stalked, base truncate, scales firm, with dark red-brown border on adaxial surface distally; apophyses slightly raised, rhombic, with strong transverse keels; umbo central, low-triangular, with slender, downcurved prickle. |
cones maturing and shed in 1–3 seasons or long-persistent, sometimes serotinous (not opening upon maturity but much later: Pinus), compound, axillary, solitary or grouped; scales overlapping, free from subtending included or exserted bracts for most of length, spirally arranged, strongly flattened, at maturity relatively thin to strongly thickened and woody (in Pinus), with 2 inverted, adaxial ovules. |
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2n | =24. |
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Pinus rigida |
Pinaceae |
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Habitat | Upland or lowland, sterile, dry to boggy soils | |||||||||||||||||||||||||
Elevation | 0–1400m (0–4600ft) | |||||||||||||||||||||||||
Distribution |
CT; DE; GA; KY; MA; MD; ME; NC; NH; NJ; NY; OH; PA; RI; SC; TN; VA; VT; WV; ON; QC
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Almost entirely in the Northern Hemisphere |
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Discussion | Pinus rigida often has poor form and is not valued highly as saw timber. It is fire successional, sprouts adventitiously, and is frequently shrubby in the northern part of its range. It is known to hybridize naturally with P. echinata. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
The Pinaceae, with a fossil record extending back to the Cretaceous (C.N. Miller Jr. 1988), constitute a clearly defined natural taxon, the basic delimiting features of which are seen in the mature seed cones: bract-scale complexes consisting of well-developed scales that are free for most of their length from the subtending bracts, two inverted ovules on the adaxial face of each scale, and usually an obvious seed wing that develops from the cone scale. The 10 genera, too, are clearly defined. The cones of certain members of the Pinaceae remain on the tree and closed for several to many years until a stimulus (often fire) causes them to open and shed their seeds. This condition, known as serotiny (adjective, serotinous), is seen in various pines (e.g., Pinus attenuata, P. banksiana, P. contorta). This primarily Northern Hemisphere family extends south to the West Indies, Central America, Japan, China, Indonesia, the Himalayas, and North Africa. The family is dominant in the vegetation of large regions including, in the flora area, forests of the boreal and Pacific regions, of the western mountains, and of the southeastern coastal plain. Only one species of the family, Pinus merkusii, crosses the equator (in Sumatra). Members of the Pinaceae are of major economic importance as producers of most of the world's softwood timber. Additionally, they are sources of pulpwood, naval stores (e.g., tar, pitch, turpentine, etc.), essential oils, and other forest products. All members of the family present in the flora, especially pines, are of varying importance to wildlife for food and cover. Many species, including most of the genera, are grown as ornamentals and shelter-belt trees and for revegetation. Most commonly seen in cultivation in the flora area are species of Abies, Cedrus, Larix, Picea, Pinus, Pseudotsuga, and Tsuga, each of these genera being represented by numerous cultivars. Keteleeria and Pseudolarix are mainly botanical garden subjects. Cathaya, the most recently described genus (1958), is apparently not yet in cultivation in North America. Among the vegetative features useful for identification of some genera of Pinaceae are the leaf scars. These are best observed on those portions of living branchlets from which leaves have fallen. Genera 10, species ca. 200 (6 genera, 66 species in the flora with 64 natives and 2 naturalized). (Discussion copyrighted by Flora of North America; reprinted with permission.) |
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Key |
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Source | FNA vol. 2. | FNA vol. 2, p. 352. | ||||||||||||||||||||||||
Parent taxa | Pinaceae > Pinus | |||||||||||||||||||||||||
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Name authority | Miller: Gard. Dict., ed. 8 Pinus no. 10. (1768) | Lindley | ||||||||||||||||||||||||
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