Pinus quadrifolia |
Pinus washoensis |
|
---|---|---|
four-needle pinyon, nut pine, Parry pine, Parry pinyon, Parry pinyon pine, piñón, piñón de California |
Washoe pine |
|
Habit | Trees to 10m; trunk to 0.5m diam., straight, much branched; crown dense, becoming rounded. | Trees to 60m; trunk to 1m diam., straight; crown pyramidal. |
Bark | red-brown, irregularly furrowed and cross-checked to irregularly rectangular, plates scaly. |
yellow-brown to reddish, fissured, plates scaly. |
Branches | spreading to ascending, persistent to trunk base; twigs slender, pale orange-brown, puberulent-glandular, aging brown to gray-brown. |
spreading-ascending; twigs stout, orangish, aging gray, rough. |
Buds | ovoid, light red-brown, ca. 0.4–0.5cm, slightly resinous. |
ovoid, red-brown, 1.5–2cm, not resinous; scale margins fringed. |
Leaves | (3–)4(–5) per fascicle, persisting 3–4 years, (2–)3–6cm × (1–)1.2–1.7mm, curved, connivent, stiff, green to blue-green, margins entire to minutely scaly-denticulate, finely serrulate, apex subulate, adaxial surfaces mostly strongly whitened with stomatal bands, abaxial surface not so but 2 subepidermal resin bands evident; sheath 0.5–0.6cm, scales soon recurved, forming rosette, shed early. |
(2–)3 per fascicle, spreading-ascending, persisting (2–)4–6(–7) years, 10–15cm × ca. 1.5mm, slightly twisted, gray-green, all surfaces with stomatal lines, margins finely serrulate, apex acuminate; sheath 1–2cm, base persistent. |
Pollen cones | ovoid, ca. 10mm, yellowish. |
cylindric, 10–20mm, red-purple. |
Seed(s) | cones maturing in 2 years, shedding seeds and falling soon thereafter, spreading, symmetric, ovoid before opening, broadly ovoid to depressed-globose when open, (3–)4–8(–10)cm, pale yellow-brown, sessile to short-stalked, apophyses thickened, strongly raised, diamond-shaped, transversely keeled, umbo subcentral, low-pyramidal or sunken, blunt. |
cones maturing in 2 years, shedding seeds soon thereafter, not persistent, spreading, slightly asymmetric, ovoid-conic before opening, broadly ovoid when open, 7–10cm, tan or pale red-brown, sessile, abaxial surface of scales darker and sharply contrasting in color with adaxial surface; apophyses slightly raised, low pyramidal; umbo central, narrowly pyramidal, tapering into short, reflexed, fine prickle. |
2n | =24. |
|
Pinus quadrifolia |
Pinus washoensis |
|
Habitat | Dry rocky sites | Dry montane forests |
Elevation | 1200–1800m (3900–5900ft) | 2100–2500m (6900–8200ft) |
Distribution |
CA; Mexico in Baja California
|
CA; NV |
Discussion | Pinus quadrifolia is the rarest pinyon in the flora. It hybridizes naturally with P. monophylla. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Pinus washoensis often occurs in large stands and resembles P. jeffreyi. The number and posture of seed-cone scales fall within the ranges given for P. jeffreyi. The abaxial surface of these scales has a significantly darker pigmentation, however; such a color contrast is not apparent in P. jeffreyi. Forest geneticists have developed hybrids between P. washoensis and related yellow pines, but no natural hybrids have been observed. Some workers regard P. washoensis as closely related to—or even conspecific with--- P. ponderosa. Of conservation concern. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 2. | FNA vol. 2. |
Parent taxa | Pinaceae > Pinus | Pinaceae > Pinus |
Sibling taxa | ||
Synonyms | P. cembroides var. parryana, P. juarezensis, P. parryana | |
Name authority | Parlatore ex Sudworth: U.S.D.A. Div. Forest. Bull. 14: 17. (1897) | H. Mason & Stockwell: Madroño 8: 62. (1945) |
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