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four-needle pinyon, nut pine, Parry pine, Parry pinyon, Parry pinyon pine, piñón, piñón de California

limber pine, pin blanc de l'ouest

Habit Trees to 10m; trunk to 0.5m diam., straight, much branched; crown dense, becoming rounded. Trees to 26m; trunk to 2m diam., straight to contorted; crown conic, becoming rounded.
Bark

red-brown, irregularly furrowed and cross-checked to irregularly rectangular, plates scaly.

gray, nearly smooth, cross-checked in age into scaly plates and ridges.

Branches

spreading to ascending, persistent to trunk base;

twigs slender, pale orange-brown, puberulent-glandular, aging brown to gray-brown.

spreading to ascending, often persistent to trunk base;

twigs pale red-brown, puberulous (rarely glabrous), slightly resinous, aging gray, smooth.

Buds

ovoid, light red-brown, ca. 0.4–0.5cm, slightly resinous.

ovoid, light red-brown, 0.9–1cm, resinous;

lower scales ciliolate along margins.

Leaves

(3–)4(–5) per fascicle, persisting 3–4 years, (2–)3–6cm × (1–)1.2–1.7mm, curved, connivent, stiff, green to blue-green, margins entire to minutely scaly-denticulate, finely serrulate, apex subulate, adaxial surfaces mostly strongly whitened with stomatal bands, abaxial surface not so but 2 subepidermal resin bands evident;

sheath 0.5–0.6cm, scales soon recurved, forming rosette, shed early.

5 per fascicle, spreading to upcurved and ascending, persisting 5–6 years, 3–7cm × 1–1.5mm, pliant, dark green, abaxial surface with less conspicuous stomatal bands than adaxial surfaces, adaxial surfaces with strong, pale stomatal bands, margins finely serrulate, apex conic-acute to acuminate;

sheath 1–1.5(–2)cm, shed early.

Pollen cones

ovoid, ca. 10mm, yellowish.

broadly ellipsoid-cylindric, ca. 15mm, pale red or yellow.

Seed(s)

cones maturing in 2 years, shedding seeds and falling soon thereafter, spreading, symmetric, ovoid before opening, broadly ovoid to depressed-globose when open, (3–)4–8(–10)cm, pale yellow-brown, sessile to short-stalked, apophyses thickened, strongly raised, diamond-shaped, transversely keeled, umbo subcentral, low-pyramidal or sunken, blunt.

cones maturing in 2 years, shedding seeds and falling soon thereafter, spreading, symmetric, lance-ovoid before opening, cylindro-ovoid when open, 7–15cm, straw-colored, resinous, sessile to short-stalked, apophyses much thickened, strongly cross-keeled, umbo terminal, depressed.

2n

=24.

Pinus quadrifolia

Pinus flexilis

Habitat Dry rocky sites High montane forests, often at timberline
Elevation 1200–1800m (3900–5900ft) (1000–)1500–3600m ((3300–)4900–11800ft)
Distribution
from FNA
CA; Mexico in Baja California
[WildflowerSearch map]
[BONAP county map]
from FNA
AZ; CA; CO; ID; MT; ND; NE; NM; NV; OR; SD; UT; WY; AB; BC
[WildflowerSearch map]
[BONAP county map]
Discussion

Pinus quadrifolia is the rarest pinyon in the flora. It hybridizes naturally with P. monophylla.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Pinus flexilis, much branched with a strongly tapering trunk, is little utilized because of its form and relative inaccessibility. It reportedly forms intermediates with P. strobiformis where the two overlap. The fresh-cut wood has the odor of turpentine.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 2. FNA vol. 2.
Parent taxa Pinaceae > Pinus Pinaceae > Pinus
Sibling taxa
P. albicaulis, P. aristata, P. attenuata, P. balfouriana, P. banksiana, P. cembroides, P. clausa, P. contorta, P. coulteri, P. echinata, P. edulis, P. elliottii, P. engelmannii, P. flexilis, P. glabra, P. jeffreyi, P. lambertiana, P. leiophylla, P. longaeva, P. monophylla, P. monticola, P. muricata, P. palustris, P. ponderosa, P. pungens, P. radiata, P. resinosa, P. rigida, P. sabiniana, P. serotina, P. strobiformis, P. strobus, P. sylvestris, P. taeda, P. torreyana, P. virginiana, P. washoensis
P. albicaulis, P. aristata, P. attenuata, P. balfouriana, P. banksiana, P. cembroides, P. clausa, P. contorta, P. coulteri, P. echinata, P. edulis, P. elliottii, P. engelmannii, P. glabra, P. jeffreyi, P. lambertiana, P. leiophylla, P. longaeva, P. monophylla, P. monticola, P. muricata, P. palustris, P. ponderosa, P. pungens, P. quadrifolia, P. radiata, P. resinosa, P. rigida, P. sabiniana, P. serotina, P. strobiformis, P. strobus, P. sylvestris, P. taeda, P. torreyana, P. virginiana, P. washoensis
Synonyms P. cembroides var. parryana, P. juarezensis, P. parryana Apinus flexilis
Name authority Parlatore ex Sudworth: U.S.D.A. Div. Forest. Bull. 14: 17. (1897) E. James: Account Exped. Pittsburgh 2: 27, 35. (1823)
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