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four-needle pinyon, nut pine, Parry pine, Parry pinyon, Parry pinyon pine, piñón, piñón de California

sand pine, scrub pine, spruce pine

Habit Trees to 10m; trunk to 0.5m diam., straight, much branched; crown dense, becoming rounded. Trees to 21m; trunk to 0.5m diam., straight and erect to leaning and crooked, much branched; crown mostly rounded or irregular.
Bark

red-brown, irregularly furrowed and cross-checked to irregularly rectangular, plates scaly.

gray to gray-brown, furrowed, with narrow, flat, irregular ridges, resin pockets absent, on upper sections of the trunk reddish to red-brown, platy becoming smooth distally.

Branches

spreading to ascending, persistent to trunk base;

twigs slender, pale orange-brown, puberulent-glandular, aging brown to gray-brown.

spreading to ascending, poorly self-pruning;

twigs slender, violet- to red-brown, rarely glaucous, aging gray, smooth.

Buds

ovoid, light red-brown, ca. 0.4–0.5cm, slightly resinous.

cylindric, purple-brown, to 1cm;

scale margins white-fringed.

Leaves

(3–)4(–5) per fascicle, persisting 3–4 years, (2–)3–6cm × (1–)1.2–1.7mm, curved, connivent, stiff, green to blue-green, margins entire to minutely scaly-denticulate, finely serrulate, apex subulate, adaxial surfaces mostly strongly whitened with stomatal bands, abaxial surface not so but 2 subepidermal resin bands evident;

sheath 0.5–0.6cm, scales soon recurved, forming rosette, shed early.

2 per fascicle, spreading-ascending, persisting 2–3 years, (3–)6–9(–10)cm × ca. 1mm, straight, slightly twisted, dark green, all surfaces with fine, inconspicuous stomatal lines, margins finely serrulate, apex short-conic;

sheath 0.3–0.5(–0.7)cm, base persistent.

Pollen cones

ovoid, ca. 10mm, yellowish.

ellipsoid, ca. 10mm, brownish yellow.

Seed(s)

cones maturing in 2 years, shedding seeds and falling soon thereafter, spreading, symmetric, ovoid before opening, broadly ovoid to depressed-globose when open, (3–)4–8(–10)cm, pale yellow-brown, sessile to short-stalked, apophyses thickened, strongly raised, diamond-shaped, transversely keeled, umbo subcentral, low-pyramidal or sunken, blunt.

cones maturing in 2 years, shedding seeds soon thereafter or often long-serotinous, long-persistent, solitary or whorled, spreading, symmetric (rarely slightly asymmetric, reflexed), lanceoloid before opening, ovoid to broadly ovoid when open, 3–8cm, red-brown, sessile or on stalks to 1cm, scales with dark red-brown, purple, or purple-gray border distally on adaxial surface;

apophyses thickened, shallowly and angulately raised, transversely rhombic, cross-keeled;

umbo central, low-pyramidal, tapering to sharp tip or weak, often deciduous prickle.

2n

=24.

Pinus quadrifolia

Pinus clausa

Habitat Dry rocky sites Fire successional in sand dunes and white sandhills
Elevation 1200–1800m (3900–5900ft) 0–60m (0–200ft)
Distribution
from FNA
CA; Mexico in Baja California
[WildflowerSearch map]
[BONAP county map]
from FNA
AL; FL
[WildflowerSearch map]
[BONAP county map]
Discussion

Pinus quadrifolia is the rarest pinyon in the flora. It hybridizes naturally with P. monophylla.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Although Pinus clausa is too profusely branched to be important for saw timber, it is managed to produce a high volume of pulpwood in northern peninsular Florida.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 2. FNA vol. 2.
Parent taxa Pinaceae > Pinus Pinaceae > Pinus
Sibling taxa
P. albicaulis, P. aristata, P. attenuata, P. balfouriana, P. banksiana, P. cembroides, P. clausa, P. contorta, P. coulteri, P. echinata, P. edulis, P. elliottii, P. engelmannii, P. flexilis, P. glabra, P. jeffreyi, P. lambertiana, P. leiophylla, P. longaeva, P. monophylla, P. monticola, P. muricata, P. palustris, P. ponderosa, P. pungens, P. radiata, P. resinosa, P. rigida, P. sabiniana, P. serotina, P. strobiformis, P. strobus, P. sylvestris, P. taeda, P. torreyana, P. virginiana, P. washoensis
P. albicaulis, P. aristata, P. attenuata, P. balfouriana, P. banksiana, P. cembroides, P. contorta, P. coulteri, P. echinata, P. edulis, P. elliottii, P. engelmannii, P. flexilis, P. glabra, P. jeffreyi, P. lambertiana, P. leiophylla, P. longaeva, P. monophylla, P. monticola, P. muricata, P. palustris, P. ponderosa, P. pungens, P. quadrifolia, P. radiata, P. resinosa, P. rigida, P. sabiniana, P. serotina, P. strobiformis, P. strobus, P. sylvestris, P. taeda, P. torreyana, P. virginiana, P. washoensis
Synonyms P. cembroides var. parryana, P. juarezensis, P. parryana P. inops var. clausa, P. clausa var. immuginata
Name authority Parlatore ex Sudworth: U.S.D.A. Div. Forest. Bull. 14: 17. (1897) (Chapman ex Engelmann) Sargent: Rep. For. N. America 199. (1884)
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