Pinus muricata |
Pinus quadrifolia |
|
---|---|---|
Bishop pine |
four-needle pinyon, nut pine, Parry pine, Parry pinyon, Parry pinyon pine, piñón, piñón de California |
|
Habit | Trees to 24m; trunk to 0.9m diam., straight to contorted; crown becoming rounded, flattened, or irregular. | Trees to 10m; trunk to 0.5m diam., straight, much branched; crown dense, becoming rounded. |
Bark | dark gray, deeply furrowed, ridges long, scaly-plated. |
red-brown, irregularly furrowed and cross-checked to irregularly rectangular, plates scaly. |
Branches | spreading-ascending, often contorted; twigs stout to slender, orange-brown, aging darker brown, rough. |
spreading to ascending, persistent to trunk base; twigs slender, pale orange-brown, puberulent-glandular, aging brown to gray-brown. |
Buds | ovoid-cylindric, dark brown, 1–2.5cm, resinous. |
ovoid, light red-brown, ca. 0.4–0.5cm, slightly resinous. |
Leaves | 2 per fascicle, spreading to upcurved, persisting 2–3 years, 8–15cm × (1.2–)1.5(–2)mm, slightly twisted, dark yellow-green, all surfaces with stomatal lines, margins strongly serrulate, apex abruptly conic-acute; sheath to 1.5cm, base persistent. |
(3–)4(–5) per fascicle, persisting 3–4 years, (2–)3–6cm × (1–)1.2–1.7mm, curved, connivent, stiff, green to blue-green, margins entire to minutely scaly-denticulate, finely serrulate, apex subulate, adaxial surfaces mostly strongly whitened with stomatal bands, abaxial surface not so but 2 subepidermal resin bands evident; sheath 0.5–0.6cm, scales soon recurved, forming rosette, shed early. |
Pollen cones | ellipsoid, to 5mm, orange. |
ovoid, ca. 10mm, yellowish. |
Seed(s) | cones maturing in 3 years, serotinous, long-persistent, mostly in whorls, mostly asymmetric, lanceoloid-ovoid before opening, curved-ovoid when open, 4–9cm, glossy bright to pale red-brown, sessile or on stalks to 1cm, mostly downcurved, scales with deep red-brown border distally on adaxial surface; apophyses much thickened, the abaxial ones progressively more angulately dome-shaped toward base of cone; umbo central, a stout-based, curved claw. |
cones maturing in 2 years, shedding seeds and falling soon thereafter, spreading, symmetric, ovoid before opening, broadly ovoid to depressed-globose when open, (3–)4–8(–10)cm, pale yellow-brown, sessile to short-stalked, apophyses thickened, strongly raised, diamond-shaped, transversely keeled, umbo subcentral, low-pyramidal or sunken, blunt. |
2n | =24. |
|
Pinus muricata |
Pinus quadrifolia |
|
Habitat | Dry ridges to coastal, windshorn forests, often in or around bogs | Dry rocky sites |
Elevation | 0–300m (0–1000ft) | 1200–1800m (3900–5900ft) |
Distribution |
CA; Mexico in Baja California
|
CA; Mexico in Baja California
|
Discussion | The several varieties described for Pinus muricata reflect the high variability in leaf characters and in degree of elaboration of apophysis and umbo in this species. The extremes can sometimes occur together. Of conservation concern. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Pinus quadrifolia is the rarest pinyon in the flora. It hybridizes naturally with P. monophylla. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 2. | FNA vol. 2. |
Parent taxa | Pinaceae > Pinus | Pinaceae > Pinus |
Sibling taxa | ||
Synonyms | P. muricata var. borealis, P. muricata var. cedrosensis, P. muricata var. stantonii, P. radiata var. binata, P. remorata | P. cembroides var. parryana, P. juarezensis, P. parryana |
Name authority | D. Don: Trans. Linn. Soc. London, Bot. 17: 441. (1836) | Parlatore ex Sudworth: U.S.D.A. Div. Forest. Bull. 14: 17. (1897) |
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