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Bishop pine

sand pine, scrub pine, spruce pine

Habit Trees to 24m; trunk to 0.9m diam., straight to contorted; crown becoming rounded, flattened, or irregular. Trees to 21m; trunk to 0.5m diam., straight and erect to leaning and crooked, much branched; crown mostly rounded or irregular.
Bark

dark gray, deeply furrowed, ridges long, scaly-plated.

gray to gray-brown, furrowed, with narrow, flat, irregular ridges, resin pockets absent, on upper sections of the trunk reddish to red-brown, platy becoming smooth distally.

Branches

spreading-ascending, often contorted;

twigs stout to slender, orange-brown, aging darker brown, rough.

spreading to ascending, poorly self-pruning;

twigs slender, violet- to red-brown, rarely glaucous, aging gray, smooth.

Buds

ovoid-cylindric, dark brown, 1–2.5cm, resinous.

cylindric, purple-brown, to 1cm;

scale margins white-fringed.

Leaves

2 per fascicle, spreading to upcurved, persisting 2–3 years, 8–15cm × (1.2–)1.5(–2)mm, slightly twisted, dark yellow-green, all surfaces with stomatal lines, margins strongly serrulate, apex abruptly conic-acute;

sheath to 1.5cm, base persistent.

2 per fascicle, spreading-ascending, persisting 2–3 years, (3–)6–9(–10)cm × ca. 1mm, straight, slightly twisted, dark green, all surfaces with fine, inconspicuous stomatal lines, margins finely serrulate, apex short-conic;

sheath 0.3–0.5(–0.7)cm, base persistent.

Pollen cones

ellipsoid, to 5mm, orange.

ellipsoid, ca. 10mm, brownish yellow.

Seed(s)

cones maturing in 3 years, serotinous, long-persistent, mostly in whorls, mostly asymmetric, lanceoloid-ovoid before opening, curved-ovoid when open, 4–9cm, glossy bright to pale red-brown, sessile or on stalks to 1cm, mostly downcurved, scales with deep red-brown border distally on adaxial surface;

apophyses much thickened, the abaxial ones progressively more angulately dome-shaped toward base of cone;

umbo central, a stout-based, curved claw.

cones maturing in 2 years, shedding seeds soon thereafter or often long-serotinous, long-persistent, solitary or whorled, spreading, symmetric (rarely slightly asymmetric, reflexed), lanceoloid before opening, ovoid to broadly ovoid when open, 3–8cm, red-brown, sessile or on stalks to 1cm, scales with dark red-brown, purple, or purple-gray border distally on adaxial surface;

apophyses thickened, shallowly and angulately raised, transversely rhombic, cross-keeled;

umbo central, low-pyramidal, tapering to sharp tip or weak, often deciduous prickle.

2n

=24.

=24.

Pinus muricata

Pinus clausa

Habitat Dry ridges to coastal, windshorn forests, often in or around bogs Fire successional in sand dunes and white sandhills
Elevation 0–300m (0–1000ft) 0–60m (0–200ft)
Distribution
from FNA
CA; Mexico in Baja California
[WildflowerSearch map]
[BONAP county map]
from FNA
AL; FL
[WildflowerSearch map]
[BONAP county map]
Discussion

The several varieties described for Pinus muricata reflect the high variability in leaf characters and in degree of elaboration of apophysis and umbo in this species. The extremes can sometimes occur together.

Of conservation concern.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Although Pinus clausa is too profusely branched to be important for saw timber, it is managed to produce a high volume of pulpwood in northern peninsular Florida.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 2. FNA vol. 2.
Parent taxa Pinaceae > Pinus Pinaceae > Pinus
Sibling taxa
P. albicaulis, P. aristata, P. attenuata, P. balfouriana, P. banksiana, P. cembroides, P. clausa, P. contorta, P. coulteri, P. echinata, P. edulis, P. elliottii, P. engelmannii, P. flexilis, P. glabra, P. jeffreyi, P. lambertiana, P. leiophylla, P. longaeva, P. monophylla, P. monticola, P. palustris, P. ponderosa, P. pungens, P. quadrifolia, P. radiata, P. resinosa, P. rigida, P. sabiniana, P. serotina, P. strobiformis, P. strobus, P. sylvestris, P. taeda, P. torreyana, P. virginiana, P. washoensis
P. albicaulis, P. aristata, P. attenuata, P. balfouriana, P. banksiana, P. cembroides, P. contorta, P. coulteri, P. echinata, P. edulis, P. elliottii, P. engelmannii, P. flexilis, P. glabra, P. jeffreyi, P. lambertiana, P. leiophylla, P. longaeva, P. monophylla, P. monticola, P. muricata, P. palustris, P. ponderosa, P. pungens, P. quadrifolia, P. radiata, P. resinosa, P. rigida, P. sabiniana, P. serotina, P. strobiformis, P. strobus, P. sylvestris, P. taeda, P. torreyana, P. virginiana, P. washoensis
Synonyms P. muricata var. borealis, P. muricata var. cedrosensis, P. muricata var. stantonii, P. radiata var. binata, P. remorata P. inops var. clausa, P. clausa var. immuginata
Name authority D. Don: Trans. Linn. Soc. London, Bot. 17: 441. (1836) (Chapman ex Engelmann) Sargent: Rep. For. N. America 199. (1884)
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