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bristlecone pine, Great Basin bristlecone pine, intermountain bristlecone pine

Chihuahua white pine, Mexican white pine, pino enano, Southwestern white pine

Habit Trees to 16m; trunk to 2m diam., strongly tapering; crown rounded, flattened (sheared), or irregular. Trees to 30m; trunk to 0.9m diam., slender, straight; crown conic, becoming rounded to irregular.
Bark

red-brown, shallowly to deeply fissured with thick, scaly, irregular, blocky ridges.

gray, aging red-brown, furrowed, with narrow, irregular, scaly ridges.

Branches

contorted, pendent;

twigs pale red-brown, aging gray to yellow-gray, puberulent, young branches resembling long bottlebrushes because of persistent leaves.

spreading-ascending;

twigs slender, pale red-brown, puberulous or glabrous, sometimes glaucous, aging gray or gray-brown, smooth.

Buds

ovoid-acuminate, pale red-brown, ca. 1cm, resinous.

ellipsoid, red-brown, ca. 1cm, resinous.

Leaves

mostly 5 per fascicle, upcurved, persisting 10–30 years, 1.5–3.5cm × 0.8–1.2mm, mostly connivent, deep yellow-green, with few resin splotches but often scurfy with pale scales, abaxial surface without median groove but with 2 subepidermal but evident resin bands, adaxial surfaces conspicuously whitened with stomates, margins entire or remotely and finely serrulate distally, apex bluntly acute to short-acuminate;

sheath ca. 1cm, soon forming rosette, shed early.

5 per fascicle, spreading to ascending-upcurved, persisting 3–5 years, 4–9cm × 0.6–1mm, straight, slightly twisted, pliant, dark green to blue-green, abaxial surface without evident stomatal lines, adaxial surfaces conspicuously whitened by narrow stomatal lines, margins sharp, razorlike and entire to finely serrulate, apex narrowly acute to short-subulate;

sheath 1.5–2cm, shed early.

Pollen cones

cylindro-ellipsoid, 7–10mm, purple-red.

cylindric, ca. 6–10mm, pale yellow-brown.

Seed(s)

cones maturing in 2 years, shedding seeds and falling soon thereafter, spreading, symmetric, lance-cylindric with rounded base before opening, lance-cylindric to narrowly ovoid when open, 6–9.5cm, purple, aging red-brown, nearly sessile;

apophyses much thickened, sharply keeled;

umbo central, raised on low buttress, truncate to umbilicate, abruptly narrowed to slender but stiff, variable prickle 1–6mm, resin exudate pale.

cones maturing in 2 years, shedding seeds and falling soon thereafter, pendent, symmetric, lance-cylindric before opening, broadly lance-cylindric when open, 15–25cm, creamy brown to light yellow-brown, stalks to 6cm;

apophyses somewhat thickened, strongly cross-keeled, tip reflexed;

umbo terminal, low.

2n

=24.

Pinus longaeva

Pinus strobiformis

Habitat Subalpine and alpine Arid to moist summit elevations, montane forests
Elevation 1700–3400m (5600–11200ft) 1900–3000m (6200–9800ft)
Distribution
from FNA
CA; NV; UT
[WildflowerSearch map]
[BONAP county map]
from FNA
AZ; NM; TX; n Mexico
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[BONAP county map]
Discussion

Pinus longaeva is considered by dendrochronologists to be the longest-lived tree. One tree was estimated to be 5000 years old.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

In the northern part of the range, Pinus strobiformis overlaps P. flexilis and reportedly hybridizes with it. On average P. strobiformis has longer, more slender leaves and thinner, more spreading-tipped apophyses than are found in P. flexilis, and stomatal bands are not evident on the abaxial surface of its leaves.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 2. FNA vol. 2.
Parent taxa Pinaceae > Pinus Pinaceae > Pinus
Sibling taxa
P. albicaulis, P. aristata, P. attenuata, P. balfouriana, P. banksiana, P. cembroides, P. clausa, P. contorta, P. coulteri, P. echinata, P. edulis, P. elliottii, P. engelmannii, P. flexilis, P. glabra, P. jeffreyi, P. lambertiana, P. leiophylla, P. monophylla, P. monticola, P. muricata, P. palustris, P. ponderosa, P. pungens, P. quadrifolia, P. radiata, P. resinosa, P. rigida, P. sabiniana, P. serotina, P. strobiformis, P. strobus, P. sylvestris, P. taeda, P. torreyana, P. virginiana, P. washoensis
P. albicaulis, P. aristata, P. attenuata, P. balfouriana, P. banksiana, P. cembroides, P. clausa, P. contorta, P. coulteri, P. echinata, P. edulis, P. elliottii, P. engelmannii, P. flexilis, P. glabra, P. jeffreyi, P. lambertiana, P. leiophylla, P. longaeva, P. monophylla, P. monticola, P. muricata, P. palustris, P. ponderosa, P. pungens, P. quadrifolia, P. radiata, P. resinosa, P. rigida, P. sabiniana, P. serotina, P. strobus, P. sylvestris, P. taeda, P. torreyana, P. virginiana, P. washoensis
Synonyms P. aristata var. longaeva P. ayacahuite var. brachyptera, P. ayacahuite var. reflexa, P. ayacahuite var. strobiformis, P. flexilis var. reflexa, P. reflexa
Name authority D. K. Bailey: Ann. Missouri Bot. Gard. 57: 243. (1970) Engelmann: in Wislizenus, Mem. Tour N. Mexico 102. (1848)
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