Pinus longaeva |
Pinus quadrifolia |
|
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bristlecone pine, Great Basin bristlecone pine, intermountain bristlecone pine |
four-needle pinyon, nut pine, Parry pine, Parry pinyon, Parry pinyon pine, piñón, piñón de California |
|
Habit | Trees to 16m; trunk to 2m diam., strongly tapering; crown rounded, flattened (sheared), or irregular. | Trees to 10m; trunk to 0.5m diam., straight, much branched; crown dense, becoming rounded. |
Bark | red-brown, shallowly to deeply fissured with thick, scaly, irregular, blocky ridges. |
red-brown, irregularly furrowed and cross-checked to irregularly rectangular, plates scaly. |
Branches | contorted, pendent; twigs pale red-brown, aging gray to yellow-gray, puberulent, young branches resembling long bottlebrushes because of persistent leaves. |
spreading to ascending, persistent to trunk base; twigs slender, pale orange-brown, puberulent-glandular, aging brown to gray-brown. |
Buds | ovoid-acuminate, pale red-brown, ca. 1cm, resinous. |
ovoid, light red-brown, ca. 0.4–0.5cm, slightly resinous. |
Leaves | mostly 5 per fascicle, upcurved, persisting 10–30 years, 1.5–3.5cm × 0.8–1.2mm, mostly connivent, deep yellow-green, with few resin splotches but often scurfy with pale scales, abaxial surface without median groove but with 2 subepidermal but evident resin bands, adaxial surfaces conspicuously whitened with stomates, margins entire or remotely and finely serrulate distally, apex bluntly acute to short-acuminate; sheath ca. 1cm, soon forming rosette, shed early. |
(3–)4(–5) per fascicle, persisting 3–4 years, (2–)3–6cm × (1–)1.2–1.7mm, curved, connivent, stiff, green to blue-green, margins entire to minutely scaly-denticulate, finely serrulate, apex subulate, adaxial surfaces mostly strongly whitened with stomatal bands, abaxial surface not so but 2 subepidermal resin bands evident; sheath 0.5–0.6cm, scales soon recurved, forming rosette, shed early. |
Pollen cones | cylindro-ellipsoid, 7–10mm, purple-red. |
ovoid, ca. 10mm, yellowish. |
Seed(s) | cones maturing in 2 years, shedding seeds and falling soon thereafter, spreading, symmetric, lance-cylindric with rounded base before opening, lance-cylindric to narrowly ovoid when open, 6–9.5cm, purple, aging red-brown, nearly sessile; apophyses much thickened, sharply keeled; umbo central, raised on low buttress, truncate to umbilicate, abruptly narrowed to slender but stiff, variable prickle 1–6mm, resin exudate pale. |
cones maturing in 2 years, shedding seeds and falling soon thereafter, spreading, symmetric, ovoid before opening, broadly ovoid to depressed-globose when open, (3–)4–8(–10)cm, pale yellow-brown, sessile to short-stalked, apophyses thickened, strongly raised, diamond-shaped, transversely keeled, umbo subcentral, low-pyramidal or sunken, blunt. |
Pinus longaeva |
Pinus quadrifolia |
|
Habitat | Subalpine and alpine | Dry rocky sites |
Elevation | 1700–3400m (5600–11200ft) | 1200–1800m (3900–5900ft) |
Distribution |
CA; NV; UT
|
CA; Mexico in Baja California
|
Discussion | Pinus longaeva is considered by dendrochronologists to be the longest-lived tree. One tree was estimated to be 5000 years old. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Pinus quadrifolia is the rarest pinyon in the flora. It hybridizes naturally with P. monophylla. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 2. | FNA vol. 2. |
Parent taxa | Pinaceae > Pinus | Pinaceae > Pinus |
Sibling taxa | ||
Synonyms | P. aristata var. longaeva | P. cembroides var. parryana, P. juarezensis, P. parryana |
Name authority | D. K. Bailey: Ann. Missouri Bot. Gard. 57: 243. (1970) | Parlatore ex Sudworth: U.S.D.A. Div. Forest. Bull. 14: 17. (1897) |
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