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bristlecone pine, Great Basin bristlecone pine, intermountain bristlecone pine

sugar pine

Habit Trees to 16m; trunk to 2m diam., strongly tapering; crown rounded, flattened (sheared), or irregular. Trees to 75m; trunk to 3.3m diam., massive, straight; crown narrowly conic, becoming rounded.
Bark

red-brown, shallowly to deeply fissured with thick, scaly, irregular, blocky ridges.

cinnamon- to gray-brown, deeply furrowed, plates long, scaly.

Branches

contorted, pendent;

twigs pale red-brown, aging gray to yellow-gray, puberulent, young branches resembling long bottlebrushes because of persistent leaves.

spreading, distal branches ascending;

twigs gray-green to red-tan, aging gray, mostly puberulent.

Buds

ovoid-acuminate, pale red-brown, ca. 1cm, resinous.

cylindro-ovoid, red-brown, to 0.8cm, resinous.

Leaves

mostly 5 per fascicle, upcurved, persisting 10–30 years, 1.5–3.5cm × 0.8–1.2mm, mostly connivent, deep yellow-green, with few resin splotches but often scurfy with pale scales, abaxial surface without median groove but with 2 subepidermal but evident resin bands, adaxial surfaces conspicuously whitened with stomates, margins entire or remotely and finely serrulate distally, apex bluntly acute to short-acuminate;

sheath ca. 1cm, soon forming rosette, shed early.

5 per fascicle, spreading to ascending, persisting 2–4 years, 5–10cm × (0.9–)1–1.5(–2)mm, straight, slightly twisted, pliant, blue-green, abaxial surface with only a few lines evident, adaxial surfaces with evident white stomatal lines, margins finely serrulate, apex acuminate;

sheath (1–)1.5–2cm, shed early.

Pollen cones

cylindro-ellipsoid, 7–10mm, purple-red.

ellipsoid-cylindric, to 15mm, yellow.

Seed(s)

cones maturing in 2 years, shedding seeds and falling soon thereafter, spreading, symmetric, lance-cylindric with rounded base before opening, lance-cylindric to narrowly ovoid when open, 6–9.5cm, purple, aging red-brown, nearly sessile;

apophyses much thickened, sharply keeled;

umbo central, raised on low buttress, truncate to umbilicate, abruptly narrowed to slender but stiff, variable prickle 1–6mm, resin exudate pale.

cones maturing in 2 years, shedding seeds and falling soon thereafter, often clustered, pendent, symmetric, cylindric before opening, lance-cylindric to ellipsoid-cylindric when open, 25–50cm, yellow-brown, stalks 6–15cm;

apophyses somewhat thickened;

umbo terminal, depressed, resinous, slightly excurved.

2n

=24.

Pinus longaeva

Pinus lambertiana

Habitat Subalpine and alpine Montane dry to moist forests
Elevation 1700–3400m (5600–11200ft) 330–3200m (1100–10500ft)
Distribution
from FNA
CA; NV; UT
[WildflowerSearch map]
[BONAP county map]
from FNA
CA; NV; OR; Mexico in n Baja California
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[BONAP county map]
Discussion

Pinus longaeva is considered by dendrochronologists to be the longest-lived tree. One tree was estimated to be 5000 years old.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

The largest species of the genus, Pinus lambertiana also has the longest seed cone in the genus. It is an important timber tree with harvest far exceeding regrowth. It is easily distinguished from P. monticola and P. strobus by its larger cones and thicker cone scales with larger seeds; it is somewhat less reliably distinguished by its leaves, which are slightly wider and more tapering-tipped and have some stomatal lines evident on the abaxial surfaces (the lines not evident in P. monticola and P. strobus). A "sugary" resin high in cyclitols exudes from the sweet-scented fresh-cut wood.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 2. FNA vol. 2.
Parent taxa Pinaceae > Pinus Pinaceae > Pinus
Sibling taxa
P. albicaulis, P. aristata, P. attenuata, P. balfouriana, P. banksiana, P. cembroides, P. clausa, P. contorta, P. coulteri, P. echinata, P. edulis, P. elliottii, P. engelmannii, P. flexilis, P. glabra, P. jeffreyi, P. lambertiana, P. leiophylla, P. monophylla, P. monticola, P. muricata, P. palustris, P. ponderosa, P. pungens, P. quadrifolia, P. radiata, P. resinosa, P. rigida, P. sabiniana, P. serotina, P. strobiformis, P. strobus, P. sylvestris, P. taeda, P. torreyana, P. virginiana, P. washoensis
P. albicaulis, P. aristata, P. attenuata, P. balfouriana, P. banksiana, P. cembroides, P. clausa, P. contorta, P. coulteri, P. echinata, P. edulis, P. elliottii, P. engelmannii, P. flexilis, P. glabra, P. jeffreyi, P. leiophylla, P. longaeva, P. monophylla, P. monticola, P. muricata, P. palustris, P. ponderosa, P. pungens, P. quadrifolia, P. radiata, P. resinosa, P. rigida, P. sabiniana, P. serotina, P. strobiformis, P. strobus, P. sylvestris, P. taeda, P. torreyana, P. virginiana, P. washoensis
Synonyms P. aristata var. longaeva
Name authority D. K. Bailey: Ann. Missouri Bot. Gard. 57: 243. (1970) Douglas: Trans. Linn. Soc. London, Bot. 15: 500. (1827)
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