Pinus longaeva |
Pinus engelmannii |
|
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bristlecone pine, Great Basin bristlecone pine, intermountain bristlecone pine |
Apache pine, Arizona longleaf pine, pino real |
|
Habit | Trees to 16m; trunk to 2m diam., strongly tapering; crown rounded, flattened (sheared), or irregular. | Trees to 35m; trunk to 0.6m diam., straight; crown irregularly rounded, rather thin. |
Bark | red-brown, shallowly to deeply fissured with thick, scaly, irregular, blocky ridges. |
dark brown, at maturity deeply furrowed, ridges becoming yellowish, of narrow, elongate, scaly plates. |
Branches | contorted, pendent; twigs pale red-brown, aging gray to yellow-gray, puberulent, young branches resembling long bottlebrushes because of persistent leaves. |
straight to ascending; twigs stout (1–2cm thick), pale gray-brown, aging darker brown, rough. |
Buds | ovoid-acuminate, pale red-brown, ca. 1cm, resinous. |
ovoid-conic, to 2cm, resinous; scale margins pale fringed. |
Leaves | mostly 5 per fascicle, upcurved, persisting 10–30 years, 1.5–3.5cm × 0.8–1.2mm, mostly connivent, deep yellow-green, with few resin splotches but often scurfy with pale scales, abaxial surface without median groove but with 2 subepidermal but evident resin bands, adaxial surfaces conspicuously whitened with stomates, margins entire or remotely and finely serrulate distally, apex bluntly acute to short-acuminate; sheath ca. 1cm, soon forming rosette, shed early. |
3(–5) per fascicle, spreading-ascending, often drooping, forming a brush at twig tips, persisting 2 years, (20–)25–45cm × 2mm, dull green, all surfaces with fine stomatal lines, margins coarsely serrulate, apex conic-subulate; sheath 3–4cm, base persistent. |
Pollen cones | cylindro-ellipsoid, 7–10mm, purple-red. |
cylindric, ca. 25mm, yellow to yellow-brown. |
Seed(s) | cones maturing in 2 years, shedding seeds and falling soon thereafter, spreading, symmetric, lance-cylindric with rounded base before opening, lance-cylindric to narrowly ovoid when open, 6–9.5cm, purple, aging red-brown, nearly sessile; apophyses much thickened, sharply keeled; umbo central, raised on low buttress, truncate to umbilicate, abruptly narrowed to slender but stiff, variable prickle 1–6mm, resin exudate pale. |
cones maturing in 2 years and shedding seeds soon thereafter, not persistent, terminal, sometimes curved, often asymmetric, lance-ovoid before opening, ovoid when open, 11–14cm, light dull brown, nearly sessile or short-stalked; apophyses rhombic, somewhat to quite elongate, strongly raised toward outer cone base, sometimes curved, strongly cross-keeled, narrowed to thick, curved, broadly triangular-based umbo, this often producing outcurved claw. |
2n | =24. |
|
Pinus longaeva |
Pinus engelmannii |
|
Habitat | Subalpine and alpine | High and dry mountain ranges, valleys, and plateaus |
Elevation | 1700–3400m (5600–11200ft) | 1500–2500m (4900–8200ft) |
Distribution |
CA; NV; UT
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AZ; NM; Mexico
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Discussion | Pinus longaeva is considered by dendrochronologists to be the longest-lived tree. One tree was estimated to be 5000 years old. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
In general appearance Pinus engelmannii much resembles P. palustris with its short-persistent, long leaves (but in this species drooping) and in its tendency to form a grass stage. It has a deep taproot as do P. palustris and P. ponderosa. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 2. | FNA vol. 2. |
Parent taxa | Pinaceae > Pinus | Pinaceae > Pinus |
Sibling taxa | ||
Synonyms | P. aristata var. longaeva | P. macrophylla, P. apacheca, P. latifolia |
Name authority | D. K. Bailey: Ann. Missouri Bot. Gard. 57: 243. (1970) | Carrière |
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