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bristlecone pine, Great Basin bristlecone pine, intermountain bristlecone pine

Apache pine, Arizona longleaf pine, pino real

Habit Trees to 16m; trunk to 2m diam., strongly tapering; crown rounded, flattened (sheared), or irregular. Trees to 35m; trunk to 0.6m diam., straight; crown irregularly rounded, rather thin.
Bark

red-brown, shallowly to deeply fissured with thick, scaly, irregular, blocky ridges.

dark brown, at maturity deeply furrowed, ridges becoming yellowish, of narrow, elongate, scaly plates.

Branches

contorted, pendent;

twigs pale red-brown, aging gray to yellow-gray, puberulent, young branches resembling long bottlebrushes because of persistent leaves.

straight to ascending;

twigs stout (1–2cm thick), pale gray-brown, aging darker brown, rough.

Buds

ovoid-acuminate, pale red-brown, ca. 1cm, resinous.

ovoid-conic, to 2cm, resinous;

scale margins pale fringed.

Leaves

mostly 5 per fascicle, upcurved, persisting 10–30 years, 1.5–3.5cm × 0.8–1.2mm, mostly connivent, deep yellow-green, with few resin splotches but often scurfy with pale scales, abaxial surface without median groove but with 2 subepidermal but evident resin bands, adaxial surfaces conspicuously whitened with stomates, margins entire or remotely and finely serrulate distally, apex bluntly acute to short-acuminate;

sheath ca. 1cm, soon forming rosette, shed early.

3(–5) per fascicle, spreading-ascending, often drooping, forming a brush at twig tips, persisting 2 years, (20–)25–45cm × 2mm, dull green, all surfaces with fine stomatal lines, margins coarsely serrulate, apex conic-subulate;

sheath 3–4cm, base persistent.

Pollen cones

cylindro-ellipsoid, 7–10mm, purple-red.

cylindric, ca. 25mm, yellow to yellow-brown.

Seed(s)

cones maturing in 2 years, shedding seeds and falling soon thereafter, spreading, symmetric, lance-cylindric with rounded base before opening, lance-cylindric to narrowly ovoid when open, 6–9.5cm, purple, aging red-brown, nearly sessile;

apophyses much thickened, sharply keeled;

umbo central, raised on low buttress, truncate to umbilicate, abruptly narrowed to slender but stiff, variable prickle 1–6mm, resin exudate pale.

cones maturing in 2 years and shedding seeds soon thereafter, not persistent, terminal, sometimes curved, often asymmetric, lance-ovoid before opening, ovoid when open, 11–14cm, light dull brown, nearly sessile or short-stalked;

apophyses rhombic, somewhat to quite elongate, strongly raised toward outer cone base, sometimes curved, strongly cross-keeled, narrowed to thick, curved, broadly triangular-based umbo, this often producing outcurved claw.

2n

=24.

Pinus longaeva

Pinus engelmannii

Habitat Subalpine and alpine High and dry mountain ranges, valleys, and plateaus
Elevation 1700–3400m (5600–11200ft) 1500–2500m (4900–8200ft)
Distribution
from FNA
CA; NV; UT
[WildflowerSearch map]
[BONAP county map]
from FNA
AZ; NM; Mexico
[WildflowerSearch map]
[BONAP county map]
Discussion

Pinus longaeva is considered by dendrochronologists to be the longest-lived tree. One tree was estimated to be 5000 years old.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

In general appearance Pinus engelmannii much resembles P. palustris with its short-persistent, long leaves (but in this species drooping) and in its tendency to form a grass stage. It has a deep taproot as do P. palustris and P. ponderosa.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 2. FNA vol. 2.
Parent taxa Pinaceae > Pinus Pinaceae > Pinus
Sibling taxa
P. albicaulis, P. aristata, P. attenuata, P. balfouriana, P. banksiana, P. cembroides, P. clausa, P. contorta, P. coulteri, P. echinata, P. edulis, P. elliottii, P. engelmannii, P. flexilis, P. glabra, P. jeffreyi, P. lambertiana, P. leiophylla, P. monophylla, P. monticola, P. muricata, P. palustris, P. ponderosa, P. pungens, P. quadrifolia, P. radiata, P. resinosa, P. rigida, P. sabiniana, P. serotina, P. strobiformis, P. strobus, P. sylvestris, P. taeda, P. torreyana, P. virginiana, P. washoensis
P. albicaulis, P. aristata, P. attenuata, P. balfouriana, P. banksiana, P. cembroides, P. clausa, P. contorta, P. coulteri, P. echinata, P. edulis, P. elliottii, P. flexilis, P. glabra, P. jeffreyi, P. lambertiana, P. leiophylla, P. longaeva, P. monophylla, P. monticola, P. muricata, P. palustris, P. ponderosa, P. pungens, P. quadrifolia, P. radiata, P. resinosa, P. rigida, P. sabiniana, P. serotina, P. strobiformis, P. strobus, P. sylvestris, P. taeda, P. torreyana, P. virginiana, P. washoensis
Synonyms P. aristata var. longaeva P. macrophylla, P. apacheca, P. latifolia
Name authority D. K. Bailey: Ann. Missouri Bot. Gard. 57: 243. (1970) Carrière
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