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bristlecone pine, Great Basin bristlecone pine, intermountain bristlecone pine

big-cone pine, Coulter pine

Habit Trees to 16m; trunk to 2m diam., strongly tapering; crown rounded, flattened (sheared), or irregular. Trees to 24m; trunk to 1m diam., straight to contorted; crown broad, thin, irregular.
Bark

red-brown, shallowly to deeply fissured with thick, scaly, irregular, blocky ridges.

dark gray-brown to near black, deeply furrowed, with long, scaly, irregularly anastomosing, rounded ridges.

Branches

contorted, pendent;

twigs pale red-brown, aging gray to yellow-gray, puberulent, young branches resembling long bottlebrushes because of persistent leaves.

often ascending;

twigs stout to moderately slender, violet-brown, often glaucous, aging gray-brown, rough.

Buds

ovoid-acuminate, pale red-brown, ca. 1cm, resinous.

ovoid, deep red-brown, 1.5(–3)cm, resinous;

scale margins white-fringed, apex cuspidate.

Leaves

mostly 5 per fascicle, upcurved, persisting 10–30 years, 1.5–3.5cm × 0.8–1.2mm, mostly connivent, deep yellow-green, with few resin splotches but often scurfy with pale scales, abaxial surface without median groove but with 2 subepidermal but evident resin bands, adaxial surfaces conspicuously whitened with stomates, margins entire or remotely and finely serrulate distally, apex bluntly acute to short-acuminate;

sheath ca. 1cm, soon forming rosette, shed early.

3 per fascicle, slightly spreading, not drooping, mostly ascending in a brush, persisting 3–4 years, 15–30cm × ca. 2mm, slightly curved or straight, twisted, dusty gray-green, all surfaces with pale, fine stomatal lines, margins serrulate, apex abruptly subulate;

sheath 2–4cm, base persistent.

Pollen cones

cylindro-ellipsoid, 7–10mm, purple-red.

ovoid to cylindric, to 25mm, light purple-brown, aging orange-brown.

Seed(s)

cones maturing in 2 years, shedding seeds and falling soon thereafter, spreading, symmetric, lance-cylindric with rounded base before opening, lance-cylindric to narrowly ovoid when open, 6–9.5cm, purple, aging red-brown, nearly sessile;

apophyses much thickened, sharply keeled;

umbo central, raised on low buttress, truncate to umbilicate, abruptly narrowed to slender but stiff, variable prickle 1–6mm, resin exudate pale.

cones maturing in 2 years, gradually shedding seeds thereafter and moderately persistent, massive, heavy, drooping, asymmetric at base, narrowly ovoid before opening, ovoid-cylindric when open, 20–35cm, pale yellow-brown, resinous, stalks to 3cm;

apophyses transverse-rhombic, strongly and sharply cross-keeled, elongate, curved, continuous with umbos to form long, upcurved claws 2.5–3cm.

2n

=24.

Pinus longaeva

Pinus coulteri

Habitat Subalpine and alpine Dry rocky slopes, flats, ridges, and chaparral, transitional to oak-pine woodland
Elevation 1700–3400m (5600–11200ft) 300–2100m (1000–6900ft)
Distribution
from FNA
CA; NV; UT
[WildflowerSearch map]
[BONAP county map]
from FNA
CA; Mexico in Baja California
[WildflowerSearch map]
[BONAP county map]
Discussion

Pinus longaeva is considered by dendrochronologists to be the longest-lived tree. One tree was estimated to be 5000 years old.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Pinus coulteri is the heaviest-coned pine; one who seeks its shade should wear a hardhat.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 2. FNA vol. 2.
Parent taxa Pinaceae > Pinus Pinaceae > Pinus
Sibling taxa
P. albicaulis, P. aristata, P. attenuata, P. balfouriana, P. banksiana, P. cembroides, P. clausa, P. contorta, P. coulteri, P. echinata, P. edulis, P. elliottii, P. engelmannii, P. flexilis, P. glabra, P. jeffreyi, P. lambertiana, P. leiophylla, P. monophylla, P. monticola, P. muricata, P. palustris, P. ponderosa, P. pungens, P. quadrifolia, P. radiata, P. resinosa, P. rigida, P. sabiniana, P. serotina, P. strobiformis, P. strobus, P. sylvestris, P. taeda, P. torreyana, P. virginiana, P. washoensis
P. albicaulis, P. aristata, P. attenuata, P. balfouriana, P. banksiana, P. cembroides, P. clausa, P. contorta, P. echinata, P. edulis, P. elliottii, P. engelmannii, P. flexilis, P. glabra, P. jeffreyi, P. lambertiana, P. leiophylla, P. longaeva, P. monophylla, P. monticola, P. muricata, P. palustris, P. ponderosa, P. pungens, P. quadrifolia, P. radiata, P. resinosa, P. rigida, P. sabiniana, P. serotina, P. strobiformis, P. strobus, P. sylvestris, P. taeda, P. torreyana, P. virginiana, P. washoensis
Synonyms P. aristata var. longaeva
Name authority D. K. Bailey: Ann. Missouri Bot. Gard. 57: 243. (1970) D. Don: Trans. Linn. Soc. London, Bot. 17: 440. (1836)
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