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bristlecone pine, Great Basin bristlecone pine, intermountain bristlecone pine

sand pine, scrub pine, spruce pine

Habit Trees to 16m; trunk to 2m diam., strongly tapering; crown rounded, flattened (sheared), or irregular. Trees to 21m; trunk to 0.5m diam., straight and erect to leaning and crooked, much branched; crown mostly rounded or irregular.
Bark

red-brown, shallowly to deeply fissured with thick, scaly, irregular, blocky ridges.

gray to gray-brown, furrowed, with narrow, flat, irregular ridges, resin pockets absent, on upper sections of the trunk reddish to red-brown, platy becoming smooth distally.

Branches

contorted, pendent;

twigs pale red-brown, aging gray to yellow-gray, puberulent, young branches resembling long bottlebrushes because of persistent leaves.

spreading to ascending, poorly self-pruning;

twigs slender, violet- to red-brown, rarely glaucous, aging gray, smooth.

Buds

ovoid-acuminate, pale red-brown, ca. 1cm, resinous.

cylindric, purple-brown, to 1cm;

scale margins white-fringed.

Leaves

mostly 5 per fascicle, upcurved, persisting 10–30 years, 1.5–3.5cm × 0.8–1.2mm, mostly connivent, deep yellow-green, with few resin splotches but often scurfy with pale scales, abaxial surface without median groove but with 2 subepidermal but evident resin bands, adaxial surfaces conspicuously whitened with stomates, margins entire or remotely and finely serrulate distally, apex bluntly acute to short-acuminate;

sheath ca. 1cm, soon forming rosette, shed early.

2 per fascicle, spreading-ascending, persisting 2–3 years, (3–)6–9(–10)cm × ca. 1mm, straight, slightly twisted, dark green, all surfaces with fine, inconspicuous stomatal lines, margins finely serrulate, apex short-conic;

sheath 0.3–0.5(–0.7)cm, base persistent.

Pollen cones

cylindro-ellipsoid, 7–10mm, purple-red.

ellipsoid, ca. 10mm, brownish yellow.

Seed(s)

cones maturing in 2 years, shedding seeds and falling soon thereafter, spreading, symmetric, lance-cylindric with rounded base before opening, lance-cylindric to narrowly ovoid when open, 6–9.5cm, purple, aging red-brown, nearly sessile;

apophyses much thickened, sharply keeled;

umbo central, raised on low buttress, truncate to umbilicate, abruptly narrowed to slender but stiff, variable prickle 1–6mm, resin exudate pale.

cones maturing in 2 years, shedding seeds soon thereafter or often long-serotinous, long-persistent, solitary or whorled, spreading, symmetric (rarely slightly asymmetric, reflexed), lanceoloid before opening, ovoid to broadly ovoid when open, 3–8cm, red-brown, sessile or on stalks to 1cm, scales with dark red-brown, purple, or purple-gray border distally on adaxial surface;

apophyses thickened, shallowly and angulately raised, transversely rhombic, cross-keeled;

umbo central, low-pyramidal, tapering to sharp tip or weak, often deciduous prickle.

2n

=24.

Pinus longaeva

Pinus clausa

Habitat Subalpine and alpine Fire successional in sand dunes and white sandhills
Elevation 1700–3400m (5600–11200ft) 0–60m (0–200ft)
Distribution
from FNA
CA; NV; UT
[WildflowerSearch map]
[BONAP county map]
from FNA
AL; FL
[WildflowerSearch map]
[BONAP county map]
Discussion

Pinus longaeva is considered by dendrochronologists to be the longest-lived tree. One tree was estimated to be 5000 years old.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Although Pinus clausa is too profusely branched to be important for saw timber, it is managed to produce a high volume of pulpwood in northern peninsular Florida.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 2. FNA vol. 2.
Parent taxa Pinaceae > Pinus Pinaceae > Pinus
Sibling taxa
P. albicaulis, P. aristata, P. attenuata, P. balfouriana, P. banksiana, P. cembroides, P. clausa, P. contorta, P. coulteri, P. echinata, P. edulis, P. elliottii, P. engelmannii, P. flexilis, P. glabra, P. jeffreyi, P. lambertiana, P. leiophylla, P. monophylla, P. monticola, P. muricata, P. palustris, P. ponderosa, P. pungens, P. quadrifolia, P. radiata, P. resinosa, P. rigida, P. sabiniana, P. serotina, P. strobiformis, P. strobus, P. sylvestris, P. taeda, P. torreyana, P. virginiana, P. washoensis
P. albicaulis, P. aristata, P. attenuata, P. balfouriana, P. banksiana, P. cembroides, P. contorta, P. coulteri, P. echinata, P. edulis, P. elliottii, P. engelmannii, P. flexilis, P. glabra, P. jeffreyi, P. lambertiana, P. leiophylla, P. longaeva, P. monophylla, P. monticola, P. muricata, P. palustris, P. ponderosa, P. pungens, P. quadrifolia, P. radiata, P. resinosa, P. rigida, P. sabiniana, P. serotina, P. strobiformis, P. strobus, P. sylvestris, P. taeda, P. torreyana, P. virginiana, P. washoensis
Synonyms P. aristata var. longaeva P. inops var. clausa, P. clausa var. immuginata
Name authority D. K. Bailey: Ann. Missouri Bot. Gard. 57: 243. (1970) (Chapman ex Engelmann) Sargent: Rep. For. N. America 199. (1884)
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