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bristlecone pine, Great Basin bristlecone pine, intermountain bristlecone pine

foxtail pine

Habit Trees to 16m; trunk to 2m diam., strongly tapering; crown rounded, flattened (sheared), or irregular. Trees to 22m; trunk to 2.6m diam., erect or leaning; crown broadly conic to irregular.
Bark

red-brown, shallowly to deeply fissured with thick, scaly, irregular, blocky ridges.

gray to salmon or cinnamon, platy or irregularly deep-fissured or with irregular blocky plates.

Branches

contorted, pendent;

twigs pale red-brown, aging gray to yellow-gray, puberulent, young branches resembling long bottlebrushes because of persistent leaves.

contorted, ascending to descending;

twigs red-brown, aging gray to drab yellow-gray, glabrous or puberulent, young branches resembling long bottlebrushes because of persistent leaves.

Buds

ovoid-acuminate, pale red-brown, ca. 1cm, resinous.

ovoid-acuminate, red-brown, 0.8–1cm, resinous.

Leaves

mostly 5 per fascicle, upcurved, persisting 10–30 years, 1.5–3.5cm × 0.8–1.2mm, mostly connivent, deep yellow-green, with few resin splotches but often scurfy with pale scales, abaxial surface without median groove but with 2 subepidermal but evident resin bands, adaxial surfaces conspicuously whitened with stomates, margins entire or remotely and finely serrulate distally, apex bluntly acute to short-acuminate;

sheath ca. 1cm, soon forming rosette, shed early.

5 per fascicle, upcurved, persisting 10–30 years, 1.5–4cm × 1–1.4mm, mostly connivent, deep blue- to deep yellow-green, abaxial surface without median groove but usually with 2 subepidermal but evident resin bands, adaxial surfaces conspicuously whitened by stomates, margins mostly entire to blunt, apex broadly acute to acuminate;

sheath 0.5–1cm, soon forming rosette, shed early.

Pollen cones

cylindro-ellipsoid, 7–10mm, purple-red.

ellipsoid, ca. 6–10mm, red.

Seed(s)

cones maturing in 2 years, shedding seeds and falling soon thereafter, spreading, symmetric, lance-cylindric with rounded base before opening, lance-cylindric to narrowly ovoid when open, 6–9.5cm, purple, aging red-brown, nearly sessile;

apophyses much thickened, sharply keeled;

umbo central, raised on low buttress, truncate to umbilicate, abruptly narrowed to slender but stiff, variable prickle 1–6mm, resin exudate pale.

cones maturing in 2 years, shedding seeds and falling soon thereafter, spreading, symmetric, lance-cylindric with conic base before opening, broadly lance-ovoid or ovoid to cylindric or ovoid-cylindric when open, 6–9(–11)cm, purple, aging red-brown, nearly sessile;

apophyses much thickened, rounded, larger toward cone base;

umbo central, usually depressed;

prickle absent or weak, to 1mm, resin exudates amber.

2n

=24.

Pinus longaeva

Pinus balfouriana

Habitat Subalpine and alpine Timberline and alpine meadows
Elevation 1700–3400m (5600–11200ft) 1500–3500m (4900–11500ft)
Distribution
from FNA
CA; NV; UT
[WildflowerSearch map]
[BONAP county map]
from FNA
CA
[WildflowerSearch map]
[BONAP county map]
Discussion

Pinus longaeva is considered by dendrochronologists to be the longest-lived tree. One tree was estimated to be 5000 years old.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Pinus balfouriana is the true "foxtail pine." In leaf character it is hardly, if at all, distinguishable from P. longaeva, but its strongly conic-based cones with distinctly shorter-prickled, sunken-centered umbos at once distinguish it from that species.

Plants shown to be genetically distinct from the type (differences in chemistry, form, foliage, cone orientation, and seeds) have been called Pinus balfouriana subsp. austrina R.Mastrogiuseppe & J.Mastrogiuseppe. As in several other species or species complexes in Pinus, however, there is a problem with a character gradient involving related taxa. The evidence presented by D.K. Bailey (1970) and later by R.J. Mastrogiuseppe and J.D. Mastrogiuseppe (1980) could as well be used to indicate that P. balfouriana (with its two infraspecific taxa) and P. longaeva represent a single species of three subspecies or three varieties. The more conservative view of Bailey is followed here.

Of conservation concern.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 2. FNA vol. 2.
Parent taxa Pinaceae > Pinus Pinaceae > Pinus
Sibling taxa
P. albicaulis, P. aristata, P. attenuata, P. balfouriana, P. banksiana, P. cembroides, P. clausa, P. contorta, P. coulteri, P. echinata, P. edulis, P. elliottii, P. engelmannii, P. flexilis, P. glabra, P. jeffreyi, P. lambertiana, P. leiophylla, P. monophylla, P. monticola, P. muricata, P. palustris, P. ponderosa, P. pungens, P. quadrifolia, P. radiata, P. resinosa, P. rigida, P. sabiniana, P. serotina, P. strobiformis, P. strobus, P. sylvestris, P. taeda, P. torreyana, P. virginiana, P. washoensis
P. albicaulis, P. aristata, P. attenuata, P. banksiana, P. cembroides, P. clausa, P. contorta, P. coulteri, P. echinata, P. edulis, P. elliottii, P. engelmannii, P. flexilis, P. glabra, P. jeffreyi, P. lambertiana, P. leiophylla, P. longaeva, P. monophylla, P. monticola, P. muricata, P. palustris, P. ponderosa, P. pungens, P. quadrifolia, P. radiata, P. resinosa, P. rigida, P. sabiniana, P. serotina, P. strobiformis, P. strobus, P. sylvestris, P. taeda, P. torreyana, P. virginiana, P. washoensis
Synonyms P. aristata var. longaeva P. balfouriana var. austrina, P. balfouriana subsp. austrina
Name authority D. K. Bailey: Ann. Missouri Bot. Gard. 57: 243. (1970) Greville & Balfour: in A. Murray bis, Bot. Exped. Oregon 8: no. 618, plate 3, fig. 1. (1853)
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