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Apache pine, Arizona longleaf pine, pino real

Bishop pine

Habit Trees to 35m; trunk to 0.6m diam., straight; crown irregularly rounded, rather thin. Trees to 24m; trunk to 0.9m diam., straight to contorted; crown becoming rounded, flattened, or irregular.
Bark

dark brown, at maturity deeply furrowed, ridges becoming yellowish, of narrow, elongate, scaly plates.

dark gray, deeply furrowed, ridges long, scaly-plated.

Branches

straight to ascending;

twigs stout (1–2cm thick), pale gray-brown, aging darker brown, rough.

spreading-ascending, often contorted;

twigs stout to slender, orange-brown, aging darker brown, rough.

Buds

ovoid-conic, to 2cm, resinous;

scale margins pale fringed.

ovoid-cylindric, dark brown, 1–2.5cm, resinous.

Leaves

3(–5) per fascicle, spreading-ascending, often drooping, forming a brush at twig tips, persisting 2 years, (20–)25–45cm × 2mm, dull green, all surfaces with fine stomatal lines, margins coarsely serrulate, apex conic-subulate;

sheath 3–4cm, base persistent.

2 per fascicle, spreading to upcurved, persisting 2–3 years, 8–15cm × (1.2–)1.5(–2)mm, slightly twisted, dark yellow-green, all surfaces with stomatal lines, margins strongly serrulate, apex abruptly conic-acute;

sheath to 1.5cm, base persistent.

Pollen cones

cylindric, ca. 25mm, yellow to yellow-brown.

ellipsoid, to 5mm, orange.

Seed(s)

cones maturing in 2 years and shedding seeds soon thereafter, not persistent, terminal, sometimes curved, often asymmetric, lance-ovoid before opening, ovoid when open, 11–14cm, light dull brown, nearly sessile or short-stalked;

apophyses rhombic, somewhat to quite elongate, strongly raised toward outer cone base, sometimes curved, strongly cross-keeled, narrowed to thick, curved, broadly triangular-based umbo, this often producing outcurved claw.

cones maturing in 3 years, serotinous, long-persistent, mostly in whorls, mostly asymmetric, lanceoloid-ovoid before opening, curved-ovoid when open, 4–9cm, glossy bright to pale red-brown, sessile or on stalks to 1cm, mostly downcurved, scales with deep red-brown border distally on adaxial surface;

apophyses much thickened, the abaxial ones progressively more angulately dome-shaped toward base of cone;

umbo central, a stout-based, curved claw.

2n

=24.

=24.

Pinus engelmannii

Pinus muricata

Habitat High and dry mountain ranges, valleys, and plateaus Dry ridges to coastal, windshorn forests, often in or around bogs
Elevation 1500–2500m (4900–8200ft) 0–300m (0–1000ft)
Distribution
from FNA
AZ; NM; Mexico
[WildflowerSearch map]
[BONAP county map]
from FNA
CA; Mexico in Baja California
[WildflowerSearch map]
[BONAP county map]
Discussion

In general appearance Pinus engelmannii much resembles P. palustris with its short-persistent, long leaves (but in this species drooping) and in its tendency to form a grass stage. It has a deep taproot as do P. palustris and P. ponderosa.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

The several varieties described for Pinus muricata reflect the high variability in leaf characters and in degree of elaboration of apophysis and umbo in this species. The extremes can sometimes occur together.

Of conservation concern.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 2. FNA vol. 2.
Parent taxa Pinaceae > Pinus Pinaceae > Pinus
Sibling taxa
P. albicaulis, P. aristata, P. attenuata, P. balfouriana, P. banksiana, P. cembroides, P. clausa, P. contorta, P. coulteri, P. echinata, P. edulis, P. elliottii, P. flexilis, P. glabra, P. jeffreyi, P. lambertiana, P. leiophylla, P. longaeva, P. monophylla, P. monticola, P. muricata, P. palustris, P. ponderosa, P. pungens, P. quadrifolia, P. radiata, P. resinosa, P. rigida, P. sabiniana, P. serotina, P. strobiformis, P. strobus, P. sylvestris, P. taeda, P. torreyana, P. virginiana, P. washoensis
P. albicaulis, P. aristata, P. attenuata, P. balfouriana, P. banksiana, P. cembroides, P. clausa, P. contorta, P. coulteri, P. echinata, P. edulis, P. elliottii, P. engelmannii, P. flexilis, P. glabra, P. jeffreyi, P. lambertiana, P. leiophylla, P. longaeva, P. monophylla, P. monticola, P. palustris, P. ponderosa, P. pungens, P. quadrifolia, P. radiata, P. resinosa, P. rigida, P. sabiniana, P. serotina, P. strobiformis, P. strobus, P. sylvestris, P. taeda, P. torreyana, P. virginiana, P. washoensis
Synonyms P. macrophylla, P. apacheca, P. latifolia P. muricata var. borealis, P. muricata var. cedrosensis, P. muricata var. stantonii, P. radiata var. binata, P. remorata
Name authority Carrière D. Don: Trans. Linn. Soc. London, Bot. 17: 441. (1836)
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