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Apache pine, Arizona longleaf pine, pino real

big-cone pine, Coulter pine

Habit Trees to 35m; trunk to 0.6m diam., straight; crown irregularly rounded, rather thin. Trees to 24m; trunk to 1m diam., straight to contorted; crown broad, thin, irregular.
Bark

dark brown, at maturity deeply furrowed, ridges becoming yellowish, of narrow, elongate, scaly plates.

dark gray-brown to near black, deeply furrowed, with long, scaly, irregularly anastomosing, rounded ridges.

Branches

straight to ascending;

twigs stout (1–2cm thick), pale gray-brown, aging darker brown, rough.

often ascending;

twigs stout to moderately slender, violet-brown, often glaucous, aging gray-brown, rough.

Buds

ovoid-conic, to 2cm, resinous;

scale margins pale fringed.

ovoid, deep red-brown, 1.5(–3)cm, resinous;

scale margins white-fringed, apex cuspidate.

Leaves

3(–5) per fascicle, spreading-ascending, often drooping, forming a brush at twig tips, persisting 2 years, (20–)25–45cm × 2mm, dull green, all surfaces with fine stomatal lines, margins coarsely serrulate, apex conic-subulate;

sheath 3–4cm, base persistent.

3 per fascicle, slightly spreading, not drooping, mostly ascending in a brush, persisting 3–4 years, 15–30cm × ca. 2mm, slightly curved or straight, twisted, dusty gray-green, all surfaces with pale, fine stomatal lines, margins serrulate, apex abruptly subulate;

sheath 2–4cm, base persistent.

Pollen cones

cylindric, ca. 25mm, yellow to yellow-brown.

ovoid to cylindric, to 25mm, light purple-brown, aging orange-brown.

Seed(s)

cones maturing in 2 years and shedding seeds soon thereafter, not persistent, terminal, sometimes curved, often asymmetric, lance-ovoid before opening, ovoid when open, 11–14cm, light dull brown, nearly sessile or short-stalked;

apophyses rhombic, somewhat to quite elongate, strongly raised toward outer cone base, sometimes curved, strongly cross-keeled, narrowed to thick, curved, broadly triangular-based umbo, this often producing outcurved claw.

cones maturing in 2 years, gradually shedding seeds thereafter and moderately persistent, massive, heavy, drooping, asymmetric at base, narrowly ovoid before opening, ovoid-cylindric when open, 20–35cm, pale yellow-brown, resinous, stalks to 3cm;

apophyses transverse-rhombic, strongly and sharply cross-keeled, elongate, curved, continuous with umbos to form long, upcurved claws 2.5–3cm.

2n

=24.

=24.

Pinus engelmannii

Pinus coulteri

Habitat High and dry mountain ranges, valleys, and plateaus Dry rocky slopes, flats, ridges, and chaparral, transitional to oak-pine woodland
Elevation 1500–2500m (4900–8200ft) 300–2100m (1000–6900ft)
Distribution
from FNA
AZ; NM; Mexico
[WildflowerSearch map]
[BONAP county map]
from FNA
CA; Mexico in Baja California
[WildflowerSearch map]
[BONAP county map]
Discussion

In general appearance Pinus engelmannii much resembles P. palustris with its short-persistent, long leaves (but in this species drooping) and in its tendency to form a grass stage. It has a deep taproot as do P. palustris and P. ponderosa.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Pinus coulteri is the heaviest-coned pine; one who seeks its shade should wear a hardhat.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 2. FNA vol. 2.
Parent taxa Pinaceae > Pinus Pinaceae > Pinus
Sibling taxa
P. albicaulis, P. aristata, P. attenuata, P. balfouriana, P. banksiana, P. cembroides, P. clausa, P. contorta, P. coulteri, P. echinata, P. edulis, P. elliottii, P. flexilis, P. glabra, P. jeffreyi, P. lambertiana, P. leiophylla, P. longaeva, P. monophylla, P. monticola, P. muricata, P. palustris, P. ponderosa, P. pungens, P. quadrifolia, P. radiata, P. resinosa, P. rigida, P. sabiniana, P. serotina, P. strobiformis, P. strobus, P. sylvestris, P. taeda, P. torreyana, P. virginiana, P. washoensis
P. albicaulis, P. aristata, P. attenuata, P. balfouriana, P. banksiana, P. cembroides, P. clausa, P. contorta, P. echinata, P. edulis, P. elliottii, P. engelmannii, P. flexilis, P. glabra, P. jeffreyi, P. lambertiana, P. leiophylla, P. longaeva, P. monophylla, P. monticola, P. muricata, P. palustris, P. ponderosa, P. pungens, P. quadrifolia, P. radiata, P. resinosa, P. rigida, P. sabiniana, P. serotina, P. strobiformis, P. strobus, P. sylvestris, P. taeda, P. torreyana, P. virginiana, P. washoensis
Synonyms P. macrophylla, P. apacheca, P. latifolia
Name authority Carrière D. Don: Trans. Linn. Soc. London, Bot. 17: 440. (1836)
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