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Apache pine, Arizona longleaf pine, pino real

bristlecone pine, Colorado bristlecone pine, Rocky Mountain bristlecone pine

Habit Trees to 35m; trunk to 0.6m diam., straight; crown irregularly rounded, rather thin. Trees to 15m; trunk to 1m diam., strongly tapering, twisted; crown rounded, flattened (sheared), or irregular.
Bark

dark brown, at maturity deeply furrowed, ridges becoming yellowish, of narrow, elongate, scaly plates.

gray to red-brown, shallowly fissured, with long, flat, irregular ridges.

Branches

straight to ascending;

twigs stout (1–2cm thick), pale gray-brown, aging darker brown, rough.

contorted;

twigs pale red-brown, aging gray, puberulent, young branches resembling long bottlebrushes because of persistent leaves.

Buds

ovoid-conic, to 2cm, resinous;

scale margins pale fringed.

ovoid-acuminate, pale red-brown, ca. 1cm, resinous.

Leaves

3(–5) per fascicle, spreading-ascending, often drooping, forming a brush at twig tips, persisting 2 years, (20–)25–45cm × 2mm, dull green, all surfaces with fine stomatal lines, margins coarsely serrulate, apex conic-subulate;

sheath 3–4cm, base persistent.

5 per fascicle, upcurved, persisting 10–17 years, (2–)3–4cm × 0.8–1mm, mostly connivent, deep blue-green, with drops and scales of resin, abaxial surface with strong, narrow median groove, adaxial surfaces conspicuously whitened by stomates, margins entire or distantly serrulate, apex conic-acute to conic-subulate;

sheath 0.5–1.5cm, scales soon recurving, shed early.

Pollen cones

cylindric, ca. 25mm, yellow to yellow-brown.

ellipsoid, ca. 10mm, bluish to red.

Seed(s)

cones maturing in 2 years and shedding seeds soon thereafter, not persistent, terminal, sometimes curved, often asymmetric, lance-ovoid before opening, ovoid when open, 11–14cm, light dull brown, nearly sessile or short-stalked;

apophyses rhombic, somewhat to quite elongate, strongly raised toward outer cone base, sometimes curved, strongly cross-keeled, narrowed to thick, curved, broadly triangular-based umbo, this often producing outcurved claw.

cones maturing in 2 years, shedding seeds and falling soon thereafter, spreading, symmetric, lance-cylindric before opening, lance-ovoid to ovoid or cylindric when open, 6–11cm, purple to brown, nearly sessile;

apophyses much thickened;

umbo central, with triangular base, extended into slender, brittle prickle 4–10mm.

2n

=24.

=24.

Pinus engelmannii

Pinus aristata

Habitat High and dry mountain ranges, valleys, and plateaus Subalpine and alpine
Elevation 1500–2500m (4900–8200ft) 2500–3400m (8200–11200ft)
Distribution
from FNA
AZ; NM; Mexico
[WildflowerSearch map]
[BONAP county map]
from FNA
AZ; CO; NM
[WildflowerSearch map]
[BONAP county map]
Discussion

In general appearance Pinus engelmannii much resembles P. palustris with its short-persistent, long leaves (but in this species drooping) and in its tendency to form a grass stage. It has a deep taproot as do P. palustris and P. ponderosa.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Pinus aristata has leaves usually narrower and sharper than in P. longaeva and P. balfouriana, and the leaves almost always have a narrow, median groove on the abaxial surface.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 2. FNA vol. 2.
Parent taxa Pinaceae > Pinus Pinaceae > Pinus
Sibling taxa
P. albicaulis, P. aristata, P. attenuata, P. balfouriana, P. banksiana, P. cembroides, P. clausa, P. contorta, P. coulteri, P. echinata, P. edulis, P. elliottii, P. flexilis, P. glabra, P. jeffreyi, P. lambertiana, P. leiophylla, P. longaeva, P. monophylla, P. monticola, P. muricata, P. palustris, P. ponderosa, P. pungens, P. quadrifolia, P. radiata, P. resinosa, P. rigida, P. sabiniana, P. serotina, P. strobiformis, P. strobus, P. sylvestris, P. taeda, P. torreyana, P. virginiana, P. washoensis
P. albicaulis, P. attenuata, P. balfouriana, P. banksiana, P. cembroides, P. clausa, P. contorta, P. coulteri, P. echinata, P. edulis, P. elliottii, P. engelmannii, P. flexilis, P. glabra, P. jeffreyi, P. lambertiana, P. leiophylla, P. longaeva, P. monophylla, P. monticola, P. muricata, P. palustris, P. ponderosa, P. pungens, P. quadrifolia, P. radiata, P. resinosa, P. rigida, P. sabiniana, P. serotina, P. strobiformis, P. strobus, P. sylvestris, P. taeda, P. torreyana, P. virginiana, P. washoensis
Synonyms P. macrophylla, P. apacheca, P. latifolia P. balfouriana var. aristata
Name authority Carrière Engelmann: in Parry & Engelmann, Amer. J. Sci. Arts ser. 2, 34: 331. (1862)
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