Pinus echinata
Pinus torreyana
shortleaf pine
Del Mar pine, island Torrey pine, soledad pine, Torrey pine
red-brown, scaly-plated, plates with evident resin pockets.
red-brown to purple-red, deeply furrowed with irregular, elongate, flat, scaly ridges.
spreading-ascending; 2-year-old branchlets slender (ca. 5mm or less), greenish brown to red-brown, often glaucous, aging red-brown to gray, roughened and cracking below leafy portion.
irregular, spreading-ascending, candelabralike;
twigs stout (1–2cm thick), greenish, aging deep gray-brown to near black, rough.
ovoid to cylindric, red-brown, 0.5–0.7(–1)cm, resinous.
conic-ovoid, pale brown, to 2.5cm;
scale margins white-fringed.
2(–3) per fascicle, spreading-ascending, persistent 3–5 years, (5–)7–11(–13)cm × ca. 1mm, straight, slightly twisted, gray- to yellow-green, all surfaces with fine stomatal lines, margins finely serrulate, apex abruptly acute;
sheath 0.5–1(–1.5)cm, base persistent.
mostly 5 per fascicle, ascending or spreading, persisting 3–4 years, 15–30cm × ca. 2mm, straight or curved, slightly twisted, dull gray-green, all surfaces with fine stomatal lines, margins serrulate, apex abruptly acute;
sheath to 2cm, shed early, base persistent.
cylindric, 15–20mm, yellow- to pale purple-green.
ovoid, 20–30mm, yellow.
cones maturing in 2 years, semipersistent, solitary or clustered, spreading, symmetric, lanceoloid or narrowly ovoid before opening, ovoid-conic when open, 4–6(–7)cm, red-brown, aging gray, nearly sessile or on stalks to 1cm, scales lacking contrasting dark border on adaxial surfaces distally;
umbo central, with elongate to short, stout, sharp prickle.
cones maturing in 3 years, shedding seeds soon thereafter, persisting to 5 years, lateral, massive, heavy, symmetric, ovoid before opening, broadly ovoid when open, 10–15cm, yellow- to red-brown, lustrous, stalks to 4cm;
apophyses thick, angulately dome-shaped, with 5 low convergent keels;
umbo central, forming short, curved-tipped pyramid.
=24.
Pinus echinata
Pinus torreyana
Although Pinus echinata is highly valued for timber and pulpwood, it is afflicted by root rot. It hybridizes with P. taeda, the pine most commonly associated with it.
(Discussion copyrighted by Flora of North America; reprinted with permission.)
Pinus torreyana is a rare and local Tertiary relic species whose present range is reduced to two small areas of southern California: near Del Mar (San Diego County) and on the northeastern shore of Santa Rosa Island (Santa Barbara County). Its distribution in Oligocene and Miocene (or at least that of its near ancestor) extended north to Oregon. Its harsh natural habitat elicits an unusually contorted and often sparse form, quite unlike the cleaner and taller form the species takes in cultivation.
In terms of numbers of individuals in the wild, as well as the small area occupied by natural populations, Pinus torreyana is without a doubt the rarest North American pine. As such it is under protection. Artificial crosses between it and another, more widespread Tertiary relic, P. sabiniana, have been successful.
Subspecies 2.
(Discussion copyrighted by Flora of North America; reprinted with permission.)
1. Crown of sheltered tree narrower than height of mature tree, open with well-spaced branches in natural sites; seed cones less than 13.5 cm broad, as broad as long or longer; only tip of umbo curved outward; maximum seed width ca. 12 mm. | subsp. torreyana |
1. Crown of sheltered tree broader than height of mature tree, compact in natural sites; seed cones mostly more than 13.5 cm broad, broader than long; entire umbo curved outward; maximum seed width ca. 14 mm. | subsp. insularis |
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