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shortleaf pine

four-needle pinyon, nut pine, Parry pine, Parry pinyon, Parry pinyon pine, piñón, piñón de California

Habit Trees to 40m; trunk to 1.2m diam., straight; crown rounded to conic. Trees to 10m; trunk to 0.5m diam., straight, much branched; crown dense, becoming rounded.
Bark

red-brown, scaly-plated, plates with evident resin pockets.

red-brown, irregularly furrowed and cross-checked to irregularly rectangular, plates scaly.

Branches

spreading-ascending; 2-year-old branchlets slender (ca. 5mm or less), greenish brown to red-brown, often glaucous, aging red-brown to gray, roughened and cracking below leafy portion.

spreading to ascending, persistent to trunk base;

twigs slender, pale orange-brown, puberulent-glandular, aging brown to gray-brown.

Buds

ovoid to cylindric, red-brown, 0.5–0.7(–1)cm, resinous.

ovoid, light red-brown, ca. 0.4–0.5cm, slightly resinous.

Leaves

2(–3) per fascicle, spreading-ascending, persistent 3–5 years, (5–)7–11(–13)cm × ca. 1mm, straight, slightly twisted, gray- to yellow-green, all surfaces with fine stomatal lines, margins finely serrulate, apex abruptly acute;

sheath 0.5–1(–1.5)cm, base persistent.

(3–)4(–5) per fascicle, persisting 3–4 years, (2–)3–6cm × (1–)1.2–1.7mm, curved, connivent, stiff, green to blue-green, margins entire to minutely scaly-denticulate, finely serrulate, apex subulate, adaxial surfaces mostly strongly whitened with stomatal bands, abaxial surface not so but 2 subepidermal resin bands evident;

sheath 0.5–0.6cm, scales soon recurved, forming rosette, shed early.

Pollen cones

cylindric, 15–20mm, yellow- to pale purple-green.

ovoid, ca. 10mm, yellowish.

Seed(s)

cones maturing in 2 years, semipersistent, solitary or clustered, spreading, symmetric, lanceoloid or narrowly ovoid before opening, ovoid-conic when open, 4–6(–7)cm, red-brown, aging gray, nearly sessile or on stalks to 1cm, scales lacking contrasting dark border on adaxial surfaces distally;

umbo central, with elongate to short, stout, sharp prickle.

cones maturing in 2 years, shedding seeds and falling soon thereafter, spreading, symmetric, ovoid before opening, broadly ovoid to depressed-globose when open, (3–)4–8(–10)cm, pale yellow-brown, sessile to short-stalked, apophyses thickened, strongly raised, diamond-shaped, transversely keeled, umbo subcentral, low-pyramidal or sunken, blunt.

2n

=24.

Pinus echinata

Pinus quadrifolia

Habitat Uplands, dry forests Dry rocky sites
Elevation 200–610m (700–2000ft) 1200–1800m (3900–5900ft)
Distribution
from FNA
AL; AR; DE; FL; GA; IL; KY; LA; MD; MO; MS; NC; NY; OH; OK; PA; SC; TN; TX; VA; WV
[WildflowerSearch map]
[BONAP county map]
from FNA
CA; Mexico in Baja California
[WildflowerSearch map]
[BONAP county map]
Discussion

Although Pinus echinata is highly valued for timber and pulpwood, it is afflicted by root rot. It hybridizes with P. taeda, the pine most commonly associated with it.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Pinus quadrifolia is the rarest pinyon in the flora. It hybridizes naturally with P. monophylla.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 2. FNA vol. 2.
Parent taxa Pinaceae > Pinus Pinaceae > Pinus
Sibling taxa
P. albicaulis, P. aristata, P. attenuata, P. balfouriana, P. banksiana, P. cembroides, P. clausa, P. contorta, P. coulteri, P. edulis, P. elliottii, P. engelmannii, P. flexilis, P. glabra, P. jeffreyi, P. lambertiana, P. leiophylla, P. longaeva, P. monophylla, P. monticola, P. muricata, P. palustris, P. ponderosa, P. pungens, P. quadrifolia, P. radiata, P. resinosa, P. rigida, P. sabiniana, P. serotina, P. strobiformis, P. strobus, P. sylvestris, P. taeda, P. torreyana, P. virginiana, P. washoensis
P. albicaulis, P. aristata, P. attenuata, P. balfouriana, P. banksiana, P. cembroides, P. clausa, P. contorta, P. coulteri, P. echinata, P. edulis, P. elliottii, P. engelmannii, P. flexilis, P. glabra, P. jeffreyi, P. lambertiana, P. leiophylla, P. longaeva, P. monophylla, P. monticola, P. muricata, P. palustris, P. ponderosa, P. pungens, P. radiata, P. resinosa, P. rigida, P. sabiniana, P. serotina, P. strobiformis, P. strobus, P. sylvestris, P. taeda, P. torreyana, P. virginiana, P. washoensis
Synonyms P. cembroides var. parryana, P. juarezensis, P. parryana
Name authority Miller: Gard. Dict., ed. 8 Pinus no. 12. (1768) Parlatore ex Sudworth: U.S.D.A. Div. Forest. Bull. 14: 17. (1897)
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