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shortleaf pine

big-cone pine, Coulter pine

Habit Trees to 40m; trunk to 1.2m diam., straight; crown rounded to conic. Trees to 24m; trunk to 1m diam., straight to contorted; crown broad, thin, irregular.
Bark

red-brown, scaly-plated, plates with evident resin pockets.

dark gray-brown to near black, deeply furrowed, with long, scaly, irregularly anastomosing, rounded ridges.

Branches

spreading-ascending; 2-year-old branchlets slender (ca. 5mm or less), greenish brown to red-brown, often glaucous, aging red-brown to gray, roughened and cracking below leafy portion.

often ascending;

twigs stout to moderately slender, violet-brown, often glaucous, aging gray-brown, rough.

Buds

ovoid to cylindric, red-brown, 0.5–0.7(–1)cm, resinous.

ovoid, deep red-brown, 1.5(–3)cm, resinous;

scale margins white-fringed, apex cuspidate.

Leaves

2(–3) per fascicle, spreading-ascending, persistent 3–5 years, (5–)7–11(–13)cm × ca. 1mm, straight, slightly twisted, gray- to yellow-green, all surfaces with fine stomatal lines, margins finely serrulate, apex abruptly acute;

sheath 0.5–1(–1.5)cm, base persistent.

3 per fascicle, slightly spreading, not drooping, mostly ascending in a brush, persisting 3–4 years, 15–30cm × ca. 2mm, slightly curved or straight, twisted, dusty gray-green, all surfaces with pale, fine stomatal lines, margins serrulate, apex abruptly subulate;

sheath 2–4cm, base persistent.

Pollen cones

cylindric, 15–20mm, yellow- to pale purple-green.

ovoid to cylindric, to 25mm, light purple-brown, aging orange-brown.

Seed(s)

cones maturing in 2 years, semipersistent, solitary or clustered, spreading, symmetric, lanceoloid or narrowly ovoid before opening, ovoid-conic when open, 4–6(–7)cm, red-brown, aging gray, nearly sessile or on stalks to 1cm, scales lacking contrasting dark border on adaxial surfaces distally;

umbo central, with elongate to short, stout, sharp prickle.

cones maturing in 2 years, gradually shedding seeds thereafter and moderately persistent, massive, heavy, drooping, asymmetric at base, narrowly ovoid before opening, ovoid-cylindric when open, 20–35cm, pale yellow-brown, resinous, stalks to 3cm;

apophyses transverse-rhombic, strongly and sharply cross-keeled, elongate, curved, continuous with umbos to form long, upcurved claws 2.5–3cm.

2n

=24.

=24.

Pinus echinata

Pinus coulteri

Habitat Uplands, dry forests Dry rocky slopes, flats, ridges, and chaparral, transitional to oak-pine woodland
Elevation 200–610m (700–2000ft) 300–2100m (1000–6900ft)
Distribution
from FNA
AL; AR; DE; FL; GA; IL; KY; LA; MD; MO; MS; NC; NY; OH; OK; PA; SC; TN; TX; VA; WV
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[BONAP county map]
from FNA
CA; Mexico in Baja California
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[BONAP county map]
Discussion

Although Pinus echinata is highly valued for timber and pulpwood, it is afflicted by root rot. It hybridizes with P. taeda, the pine most commonly associated with it.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Pinus coulteri is the heaviest-coned pine; one who seeks its shade should wear a hardhat.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 2. FNA vol. 2.
Parent taxa Pinaceae > Pinus Pinaceae > Pinus
Sibling taxa
P. albicaulis, P. aristata, P. attenuata, P. balfouriana, P. banksiana, P. cembroides, P. clausa, P. contorta, P. coulteri, P. edulis, P. elliottii, P. engelmannii, P. flexilis, P. glabra, P. jeffreyi, P. lambertiana, P. leiophylla, P. longaeva, P. monophylla, P. monticola, P. muricata, P. palustris, P. ponderosa, P. pungens, P. quadrifolia, P. radiata, P. resinosa, P. rigida, P. sabiniana, P. serotina, P. strobiformis, P. strobus, P. sylvestris, P. taeda, P. torreyana, P. virginiana, P. washoensis
P. albicaulis, P. aristata, P. attenuata, P. balfouriana, P. banksiana, P. cembroides, P. clausa, P. contorta, P. echinata, P. edulis, P. elliottii, P. engelmannii, P. flexilis, P. glabra, P. jeffreyi, P. lambertiana, P. leiophylla, P. longaeva, P. monophylla, P. monticola, P. muricata, P. palustris, P. ponderosa, P. pungens, P. quadrifolia, P. radiata, P. resinosa, P. rigida, P. sabiniana, P. serotina, P. strobiformis, P. strobus, P. sylvestris, P. taeda, P. torreyana, P. virginiana, P. washoensis
Name authority Miller: Gard. Dict., ed. 8 Pinus no. 12. (1768) D. Don: Trans. Linn. Soc. London, Bot. 17: 440. (1836)
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