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shortleaf pine

sand pine, scrub pine, spruce pine

Habit Trees to 40m; trunk to 1.2m diam., straight; crown rounded to conic. Trees to 21m; trunk to 0.5m diam., straight and erect to leaning and crooked, much branched; crown mostly rounded or irregular.
Bark

red-brown, scaly-plated, plates with evident resin pockets.

gray to gray-brown, furrowed, with narrow, flat, irregular ridges, resin pockets absent, on upper sections of the trunk reddish to red-brown, platy becoming smooth distally.

Branches

spreading-ascending; 2-year-old branchlets slender (ca. 5mm or less), greenish brown to red-brown, often glaucous, aging red-brown to gray, roughened and cracking below leafy portion.

spreading to ascending, poorly self-pruning;

twigs slender, violet- to red-brown, rarely glaucous, aging gray, smooth.

Buds

ovoid to cylindric, red-brown, 0.5–0.7(–1)cm, resinous.

cylindric, purple-brown, to 1cm;

scale margins white-fringed.

Leaves

2(–3) per fascicle, spreading-ascending, persistent 3–5 years, (5–)7–11(–13)cm × ca. 1mm, straight, slightly twisted, gray- to yellow-green, all surfaces with fine stomatal lines, margins finely serrulate, apex abruptly acute;

sheath 0.5–1(–1.5)cm, base persistent.

2 per fascicle, spreading-ascending, persisting 2–3 years, (3–)6–9(–10)cm × ca. 1mm, straight, slightly twisted, dark green, all surfaces with fine, inconspicuous stomatal lines, margins finely serrulate, apex short-conic;

sheath 0.3–0.5(–0.7)cm, base persistent.

Pollen cones

cylindric, 15–20mm, yellow- to pale purple-green.

ellipsoid, ca. 10mm, brownish yellow.

Seed(s)

cones maturing in 2 years, semipersistent, solitary or clustered, spreading, symmetric, lanceoloid or narrowly ovoid before opening, ovoid-conic when open, 4–6(–7)cm, red-brown, aging gray, nearly sessile or on stalks to 1cm, scales lacking contrasting dark border on adaxial surfaces distally;

umbo central, with elongate to short, stout, sharp prickle.

cones maturing in 2 years, shedding seeds soon thereafter or often long-serotinous, long-persistent, solitary or whorled, spreading, symmetric (rarely slightly asymmetric, reflexed), lanceoloid before opening, ovoid to broadly ovoid when open, 3–8cm, red-brown, sessile or on stalks to 1cm, scales with dark red-brown, purple, or purple-gray border distally on adaxial surface;

apophyses thickened, shallowly and angulately raised, transversely rhombic, cross-keeled;

umbo central, low-pyramidal, tapering to sharp tip or weak, often deciduous prickle.

2n

=24.

=24.

Pinus echinata

Pinus clausa

Habitat Uplands, dry forests Fire successional in sand dunes and white sandhills
Elevation 200–610m (700–2000ft) 0–60m (0–200ft)
Distribution
from FNA
AL; AR; DE; FL; GA; IL; KY; LA; MD; MO; MS; NC; NY; OH; OK; PA; SC; TN; TX; VA; WV
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from FNA
AL; FL
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[BONAP county map]
Discussion

Although Pinus echinata is highly valued for timber and pulpwood, it is afflicted by root rot. It hybridizes with P. taeda, the pine most commonly associated with it.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Although Pinus clausa is too profusely branched to be important for saw timber, it is managed to produce a high volume of pulpwood in northern peninsular Florida.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 2. FNA vol. 2.
Parent taxa Pinaceae > Pinus Pinaceae > Pinus
Sibling taxa
P. albicaulis, P. aristata, P. attenuata, P. balfouriana, P. banksiana, P. cembroides, P. clausa, P. contorta, P. coulteri, P. edulis, P. elliottii, P. engelmannii, P. flexilis, P. glabra, P. jeffreyi, P. lambertiana, P. leiophylla, P. longaeva, P. monophylla, P. monticola, P. muricata, P. palustris, P. ponderosa, P. pungens, P. quadrifolia, P. radiata, P. resinosa, P. rigida, P. sabiniana, P. serotina, P. strobiformis, P. strobus, P. sylvestris, P. taeda, P. torreyana, P. virginiana, P. washoensis
P. albicaulis, P. aristata, P. attenuata, P. balfouriana, P. banksiana, P. cembroides, P. contorta, P. coulteri, P. echinata, P. edulis, P. elliottii, P. engelmannii, P. flexilis, P. glabra, P. jeffreyi, P. lambertiana, P. leiophylla, P. longaeva, P. monophylla, P. monticola, P. muricata, P. palustris, P. ponderosa, P. pungens, P. quadrifolia, P. radiata, P. resinosa, P. rigida, P. sabiniana, P. serotina, P. strobiformis, P. strobus, P. sylvestris, P. taeda, P. torreyana, P. virginiana, P. washoensis
Synonyms P. inops var. clausa, P. clausa var. immuginata
Name authority Miller: Gard. Dict., ed. 8 Pinus no. 12. (1768) (Chapman ex Engelmann) Sargent: Rep. For. N. America 199. (1884)
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