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Mexican pinyon, pino piñonero, piñón

lodgepole pine, scrub pine, shore pine

Habit Shrubs or trees to 15m; trunk to 0.3m diam., strongly tapering, much branched; crown rounded. Shrubs or trees to 50m; trunk to 0.9m diam., straight to contorted; crown various according to genetic race.
Bark

red-brown to dark brown, shallowly and irregularly furrowed, ridges broad, scaly.

brown to gray- or red-brown, platy to furrowed.

Branches

spreading-ascending;

twigs red-brown, sometimes finely papillate, aging gray to gray-brown.

Twigs

slender, orange to red-brown, aging darker brown, rough.

Buds

ovoid to short cylindric, pale red-brown, 0.5–1.2cm, slightly resinous.

narrowly to broadly ovoid, dark red-brown, to 1.2cm, slightly resinous.

Leaves

(2–)3(–4) per fascicle, spreading to upcurved, persisting 3–4 years, 2–6cm × 0.6–0.9(–1)mm, connivent, 2–3-sided, blue- to gray-green, abaxial surface not conspicuously whitened with stomatal bands or if stomatal bands present, these less conspicuous than on adaxial surfaces, often with 2 subepidermal resin bands evident, adaxial surfaces conspicuously whitened with stomatal lines, margins entire to finely serrulate, apex narrowly conic or subulate;

sheath 0.5–0.7cm, scales soon recurved, forming rosette, shed early.

2 per fascicle, spreading or ascending, persisting 3–8 years, 2–8cm × 0.7–2(–3)mm, twisted, yellow-green to dark green, all surfaces with fine stomatal lines, margins finely serrulate, apex blunt to acute or narrowly acuminate;

sheath 0.3–0.6(–1)cm, persistent.

Pollen cones

ellipsoid, to 10mm, yellow.

ellipsoid to cylindric, 5–15mm, orange-red.

Seed(s)

cones maturing in 2 years, shedding seeds and falling soon thereafter, spreading, symmetric, ovoid before opening, broadly depressed-ovoid to nearly globose when open, 1–3.5cm, pale yellow- to pale red-brown, resinous, nearly sessile or short-stalked;

apophyses thickened, slightly domed, angulate, transversely keeled;

umbo subcentral, slightly raised to depressed, truncate or umbilicate.

cones maturing in 2 years or variably serotinous, variably persistent, spreading to reflexed, often curved, nearly symmetric or variably asymmetric, lanceoloid to ovoid before opening, broadly ovoid to nearly globose when open, 2–6cm, tan to pale red-brown, lustrous, nearly sessile or on stalks to 1cm;

apophyses nearly rhombic, variously elongate, cross-keeled, often mammillate toward outer cone base and on inside above middle;

umbo central, depressed-triangular, prickle barely elongate to stubby or slender and to 6mm.

Lower

branches often descending, the upper spreading or ascending.

2n

=24.

=24 (variety not indicated).

Pinus cembroides

Pinus contorta

Habitat Pinyon-juniper woodland, foothills, mesas, tablelands
Elevation 700–2300m (2300–7500ft)
Distribution
from FNA
AZ; NM; TX; Mexico
[WildflowerSearch map]
[BONAP county map]
from FNA
AK; CA; CO; ID; MT; NV; OR; SD; UT; WA; WY; AB; BC; NT; SK; YT; only in the flora
[WildflowerSearch map]
[BONAP county map]
Discussion

Pinus cembroides is the common pinyon of Mexican commerce. Populations of the Edwards Plateau, Texas, are disjunct about 150km east and north of the main area of distribution of the species, and they have been described as a distinct variety, P. cembroides var. remota Little, on the basis of thin seed shell and a higher frequency of 2-leaved fascicles in contrast to the thicker seed shell and prevalently 3-leaved fascicles in Mexican pinyon populations to the west and south. The strong overlap in nearly all character states between the populations of the Edwards Plateau and other populations makes var. remota difficult to maintain.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Pinus contorta is fire successional over most of its range and is characterized by prolific seeding and high seed viability in disturbed habitats, often resulting in extremely slow-growing, overly dense stands. Some authors consider it to consist of 4 races; these have been given various infraspecific ranks, but perhaps they are more conventionally treated as 3 varieties.

Varieties 3.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Key
1. Leaves 2-7 cm × 0.7-0.9(-1.1) mm, dark green; mature trunk with bark evidently furrowed; seed cones strongly asymmetric, strongly recurved, persistent or variously serotinous.
var. contorta
1. Leaves (4-)5-8 cm × (0.7-)1-2(-3) mm, yellow-green; mature trunk with bark not evidently furrowed; seed cones asymmetric to nearly symmetric, recurved to spreading, variously serotinous or soon shed.
→ 2
2. Seed cones asymmetric, recurved, variously serotinous, long-persistent; mid and lower apophyses mostly much domed; main branches mostly horizontally spreading, not ascending at tip.
var. latifolia
2. Seed cones nearly symmetric, mostly spreading, not serotinous, not persistent; mid and lower apophyses mostly shallowly domed; main branches ascending at tips.
var. murrayana
Source FNA vol. 2. FNA vol. 2.
Parent taxa Pinaceae > Pinus Pinaceae > Pinus
Sibling taxa
P. albicaulis, P. aristata, P. attenuata, P. balfouriana, P. banksiana, P. clausa, P. contorta, P. coulteri, P. echinata, P. edulis, P. elliottii, P. engelmannii, P. flexilis, P. glabra, P. jeffreyi, P. lambertiana, P. leiophylla, P. longaeva, P. monophylla, P. monticola, P. muricata, P. palustris, P. ponderosa, P. pungens, P. quadrifolia, P. radiata, P. resinosa, P. rigida, P. sabiniana, P. serotina, P. strobiformis, P. strobus, P. sylvestris, P. taeda, P. torreyana, P. virginiana, P. washoensis
P. albicaulis, P. aristata, P. attenuata, P. balfouriana, P. banksiana, P. cembroides, P. clausa, P. coulteri, P. echinata, P. edulis, P. elliottii, P. engelmannii, P. flexilis, P. glabra, P. jeffreyi, P. lambertiana, P. leiophylla, P. longaeva, P. monophylla, P. monticola, P. muricata, P. palustris, P. ponderosa, P. pungens, P. quadrifolia, P. radiata, P. resinosa, P. rigida, P. sabiniana, P. serotina, P. strobiformis, P. strobus, P. sylvestris, P. taeda, P. torreyana, P. virginiana, P. washoensis
Subordinate taxa
P. contorta var. contorta, P. contorta var. latifolia, P. contorta var. murrayana
Synonyms P. cembroides var. bicolor, P. cembroides var. remota, P. discolor, P. remota
Name authority Zuccarini: Abh. Math.-Phys. Cl. Königl. Bayer. Akad. Wiss. 1: 392. (1832) Douglas ex Loudon: Arbor. Frutic. Brit. 4: 2292, figs. 2210, 2211. (1838)
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