Pinus balfouriana
Pinus quadrifolia
foxtail pine
four-needle pinyon, nut pine, Parry pine, Parry pinyon, Parry pinyon pine, piñón, piñón de California
gray to salmon or cinnamon, platy or irregularly deep-fissured or with irregular blocky plates.
red-brown, irregularly furrowed and cross-checked to irregularly rectangular, plates scaly.
contorted, ascending to descending;
twigs red-brown, aging gray to drab yellow-gray, glabrous or puberulent, young branches resembling long bottlebrushes because of persistent leaves.
spreading to ascending, persistent to trunk base;
twigs slender, pale orange-brown, puberulent-glandular, aging brown to gray-brown.
ovoid-acuminate, red-brown, 0.8–1cm, resinous.
ovoid, light red-brown, ca. 0.4–0.5cm, slightly resinous.
5 per fascicle, upcurved, persisting 10–30 years, 1.5–4cm × 1–1.4mm, mostly connivent, deep blue- to deep yellow-green, abaxial surface without median groove but usually with 2 subepidermal but evident resin bands, adaxial surfaces conspicuously whitened by stomates, margins mostly entire to blunt, apex broadly acute to acuminate;
sheath 0.5–1cm, soon forming rosette, shed early.
(3–)4(–5) per fascicle, persisting 3–4 years, (2–)3–6cm × (1–)1.2–1.7mm, curved, connivent, stiff, green to blue-green, margins entire to minutely scaly-denticulate, finely serrulate, apex subulate, adaxial surfaces mostly strongly whitened with stomatal bands, abaxial surface not so but 2 subepidermal resin bands evident;
sheath 0.5–0.6cm, scales soon recurved, forming rosette, shed early.
ellipsoid, ca. 6–10mm, red.
ovoid, ca. 10mm, yellowish.
cones maturing in 2 years, shedding seeds and falling soon thereafter, spreading, symmetric, lance-cylindric with conic base before opening, broadly lance-ovoid or ovoid to cylindric or ovoid-cylindric when open, 6–9(–11)cm, purple, aging red-brown, nearly sessile;
apophyses much thickened, rounded, larger toward cone base;
umbo central, usually depressed;
prickle absent or weak, to 1mm, resin exudates amber.
cones maturing in 2 years, shedding seeds and falling soon thereafter, spreading, symmetric, ovoid before opening, broadly ovoid to depressed-globose when open, (3–)4–8(–10)cm, pale yellow-brown, sessile to short-stalked, apophyses thickened, strongly raised, diamond-shaped, transversely keeled, umbo subcentral, low-pyramidal or sunken, blunt.
=24.
Pinus balfouriana
Pinus quadrifolia
Pinus balfouriana is the true "foxtail pine." In leaf character it is hardly, if at all, distinguishable from P. longaeva, but its strongly conic-based cones with distinctly shorter-prickled, sunken-centered umbos at once distinguish it from that species.
Plants shown to be genetically distinct from the type (differences in chemistry, form, foliage, cone orientation, and seeds) have been called Pinus balfouriana subsp. austrina R.Mastrogiuseppe & J.Mastrogiuseppe. As in several other species or species complexes in Pinus, however, there is a problem with a character gradient involving related taxa. The evidence presented by D.K. Bailey (1970) and later by R.J. Mastrogiuseppe and J.D. Mastrogiuseppe (1980) could as well be used to indicate that P. balfouriana (with its two infraspecific taxa) and P. longaeva represent a single species of three subspecies or three varieties. The more conservative view of Bailey is followed here.
Of conservation concern.
(Discussion copyrighted by Flora of North America; reprinted with permission.)
Pinus quadrifolia is the rarest pinyon in the flora. It hybridizes naturally with P. monophylla.
(Discussion copyrighted by Flora of North America; reprinted with permission.)
CA
OR
Flora NW
PNW Herbaria
WildflowerSearch
iNaturalist
USDA Plants Database
