Pinus balfouriana |
Pinus aristata |
|
---|---|---|
foxtail pine |
bristlecone pine, Colorado bristlecone pine, Rocky Mountain bristlecone pine |
|
Habit | Trees to 22m; trunk to 2.6m diam., erect or leaning; crown broadly conic to irregular. | Trees to 15m; trunk to 1m diam., strongly tapering, twisted; crown rounded, flattened (sheared), or irregular. |
Bark | gray to salmon or cinnamon, platy or irregularly deep-fissured or with irregular blocky plates. |
gray to red-brown, shallowly fissured, with long, flat, irregular ridges. |
Branches | contorted, ascending to descending; twigs red-brown, aging gray to drab yellow-gray, glabrous or puberulent, young branches resembling long bottlebrushes because of persistent leaves. |
contorted; twigs pale red-brown, aging gray, puberulent, young branches resembling long bottlebrushes because of persistent leaves. |
Buds | ovoid-acuminate, red-brown, 0.8–1cm, resinous. |
ovoid-acuminate, pale red-brown, ca. 1cm, resinous. |
Leaves | 5 per fascicle, upcurved, persisting 10–30 years, 1.5–4cm × 1–1.4mm, mostly connivent, deep blue- to deep yellow-green, abaxial surface without median groove but usually with 2 subepidermal but evident resin bands, adaxial surfaces conspicuously whitened by stomates, margins mostly entire to blunt, apex broadly acute to acuminate; sheath 0.5–1cm, soon forming rosette, shed early. |
5 per fascicle, upcurved, persisting 10–17 years, (2–)3–4cm × 0.8–1mm, mostly connivent, deep blue-green, with drops and scales of resin, abaxial surface with strong, narrow median groove, adaxial surfaces conspicuously whitened by stomates, margins entire or distantly serrulate, apex conic-acute to conic-subulate; sheath 0.5–1.5cm, scales soon recurving, shed early. |
Pollen cones | ellipsoid, ca. 6–10mm, red. |
ellipsoid, ca. 10mm, bluish to red. |
Seed(s) | cones maturing in 2 years, shedding seeds and falling soon thereafter, spreading, symmetric, lance-cylindric with conic base before opening, broadly lance-ovoid or ovoid to cylindric or ovoid-cylindric when open, 6–9(–11)cm, purple, aging red-brown, nearly sessile; apophyses much thickened, rounded, larger toward cone base; umbo central, usually depressed; prickle absent or weak, to 1mm, resin exudates amber. |
cones maturing in 2 years, shedding seeds and falling soon thereafter, spreading, symmetric, lance-cylindric before opening, lance-ovoid to ovoid or cylindric when open, 6–11cm, purple to brown, nearly sessile; apophyses much thickened; umbo central, with triangular base, extended into slender, brittle prickle 4–10mm. |
2n | =24. |
=24. |
Pinus balfouriana |
Pinus aristata |
|
Habitat | Timberline and alpine meadows | Subalpine and alpine |
Elevation | 1500–3500m (4900–11500ft) | 2500–3400m (8200–11200ft) |
Distribution |
CA
|
AZ; CO; NM
|
Discussion | Pinus balfouriana is the true "foxtail pine." In leaf character it is hardly, if at all, distinguishable from P. longaeva, but its strongly conic-based cones with distinctly shorter-prickled, sunken-centered umbos at once distinguish it from that species. Plants shown to be genetically distinct from the type (differences in chemistry, form, foliage, cone orientation, and seeds) have been called Pinus balfouriana subsp. austrina R.Mastrogiuseppe & J.Mastrogiuseppe. As in several other species or species complexes in Pinus, however, there is a problem with a character gradient involving related taxa. The evidence presented by D.K. Bailey (1970) and later by R.J. Mastrogiuseppe and J.D. Mastrogiuseppe (1980) could as well be used to indicate that P. balfouriana (with its two infraspecific taxa) and P. longaeva represent a single species of three subspecies or three varieties. The more conservative view of Bailey is followed here. Of conservation concern. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Pinus aristata has leaves usually narrower and sharper than in P. longaeva and P. balfouriana, and the leaves almost always have a narrow, median groove on the abaxial surface. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 2. | FNA vol. 2. |
Parent taxa | Pinaceae > Pinus | Pinaceae > Pinus |
Sibling taxa | ||
Synonyms | P. balfouriana var. austrina, P. balfouriana subsp. austrina | P. balfouriana var. aristata |
Name authority | Greville & Balfour: in A. Murray bis, Bot. Exped. Oregon 8: no. 618, plate 3, fig. 1. (1853) | Engelmann: in Parry & Engelmann, Amer. J. Sci. Arts ser. 2, 34: 331. (1862) |
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