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knobcone pine

sugar pine

Habit Shrubs or trees to 24m; trunk to 0.8m diam., usually straight; crown mostly narrowly to broadly conic. Trees to 75m; trunk to 3.3m diam., massive, straight; crown narrowly conic, becoming rounded.
Bark

purple-brown to dark brown, shallowly and narrowly fissured, with irregular, flat, loose-scaly plates, on upper sections of trunk nearly smooth.

cinnamon- to gray-brown, deeply furrowed, plates long, scaly.

Branches

ascending;

twigs slender, red-brown.

spreading, distal branches ascending;

twigs gray-green to red-tan, aging gray, mostly puberulent.

Buds

ovoid to ovoid-cylindric, dark red-brown, aging darker, ca. 1.5cm, resinous;

scale margins fringed, apex attenuate.

cylindro-ovoid, red-brown, to 0.8cm, resinous.

Leaves

3 per fascicle, spreading or ascending, persisting 4–5 years, (8–)9–15(–20)cm × (1–)1.3–1.8mm, straight or slightly curved, twisted, yellow-green, all surfaces with fine stomatal lines, margins serrulate, apex abruptly conic-subulate;

sheath (1–)1.5–2cm, base persistent.

5 per fascicle, spreading to ascending, persisting 2–4 years, 5–10cm × (0.9–)1–1.5(–2)mm, straight, slightly twisted, pliant, blue-green, abaxial surface with only a few lines evident, adaxial surfaces with evident white stomatal lines, margins finely serrulate, apex acuminate;

sheath (1–)1.5–2cm, shed early.

Pollen cones

ellipsoid-cylindric, 10–15mm, orange-brown.

ellipsoid-cylindric, to 15mm, yellow.

Seed(s)

cones maturing in 2 years, serotinous, long-persistent, remaining closed for 20 years or more, or opening on burning, in whorls, hard and heavy, very asymmetric, lanceoloid before opening, ovoid-cylindric when open, 8–15cm, yellow- or pale red-brown, stalks to 1cm;

apophyses toward outside base increasingly elongate, mammillate or raised-angled-conic, downcurved near base, scarcely raised on branchlet side, rhombic;

umbo central, low-pyramidal, sharp, upcurved.

cones maturing in 2 years, shedding seeds and falling soon thereafter, often clustered, pendent, symmetric, cylindric before opening, lance-cylindric to ellipsoid-cylindric when open, 25–50cm, yellow-brown, stalks 6–15cm;

apophyses somewhat thickened;

umbo terminal, depressed, resinous, slightly excurved.

2n

=24.

=24.

Pinus attenuata

Pinus lambertiana

Habitat Fire successional on dry slopes and foothills of Sierra Nevada and the Cascade and Coast ranges Montane dry to moist forests
Elevation 300–1200m (1000–3900ft) 330–3200m (1100–10500ft)
Distribution
from FNA
CA; OR; Mexico in Baja California
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from FNA
CA; NV; OR; Mexico in n Baja California
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[BONAP county map]
Discussion

Pinus attenuata, mostly a chaparral species, bears cones at an early age. Its seed crops are heavy, and a hot fire permits the seeds to be released. It forms hybrids with P. muricata and P. radiata.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

The largest species of the genus, Pinus lambertiana also has the longest seed cone in the genus. It is an important timber tree with harvest far exceeding regrowth. It is easily distinguished from P. monticola and P. strobus by its larger cones and thicker cone scales with larger seeds; it is somewhat less reliably distinguished by its leaves, which are slightly wider and more tapering-tipped and have some stomatal lines evident on the abaxial surfaces (the lines not evident in P. monticola and P. strobus). A "sugary" resin high in cyclitols exudes from the sweet-scented fresh-cut wood.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 2. FNA vol. 2.
Parent taxa Pinaceae > Pinus Pinaceae > Pinus
Sibling taxa
P. albicaulis, P. aristata, P. balfouriana, P. banksiana, P. cembroides, P. clausa, P. contorta, P. coulteri, P. echinata, P. edulis, P. elliottii, P. engelmannii, P. flexilis, P. glabra, P. jeffreyi, P. lambertiana, P. leiophylla, P. longaeva, P. monophylla, P. monticola, P. muricata, P. palustris, P. ponderosa, P. pungens, P. quadrifolia, P. radiata, P. resinosa, P. rigida, P. sabiniana, P. serotina, P. strobiformis, P. strobus, P. sylvestris, P. taeda, P. torreyana, P. virginiana, P. washoensis
P. albicaulis, P. aristata, P. attenuata, P. balfouriana, P. banksiana, P. cembroides, P. clausa, P. contorta, P. coulteri, P. echinata, P. edulis, P. elliottii, P. engelmannii, P. flexilis, P. glabra, P. jeffreyi, P. leiophylla, P. longaeva, P. monophylla, P. monticola, P. muricata, P. palustris, P. ponderosa, P. pungens, P. quadrifolia, P. radiata, P. resinosa, P. rigida, P. sabiniana, P. serotina, P. strobiformis, P. strobus, P. sylvestris, P. taeda, P. torreyana, P. virginiana, P. washoensis
Synonyms P. tuberculata
Name authority Lemmon: Mining Sci. Press 64: 45. (1892) Douglas: Trans. Linn. Soc. London, Bot. 15: 500. (1827)
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