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knobcone pine

limber pine, pin blanc de l'ouest

Habit Shrubs or trees to 24m; trunk to 0.8m diam., usually straight; crown mostly narrowly to broadly conic. Trees to 26m; trunk to 2m diam., straight to contorted; crown conic, becoming rounded.
Bark

purple-brown to dark brown, shallowly and narrowly fissured, with irregular, flat, loose-scaly plates, on upper sections of trunk nearly smooth.

gray, nearly smooth, cross-checked in age into scaly plates and ridges.

Branches

ascending;

twigs slender, red-brown.

spreading to ascending, often persistent to trunk base;

twigs pale red-brown, puberulous (rarely glabrous), slightly resinous, aging gray, smooth.

Buds

ovoid to ovoid-cylindric, dark red-brown, aging darker, ca. 1.5cm, resinous;

scale margins fringed, apex attenuate.

ovoid, light red-brown, 0.9–1cm, resinous;

lower scales ciliolate along margins.

Leaves

3 per fascicle, spreading or ascending, persisting 4–5 years, (8–)9–15(–20)cm × (1–)1.3–1.8mm, straight or slightly curved, twisted, yellow-green, all surfaces with fine stomatal lines, margins serrulate, apex abruptly conic-subulate;

sheath (1–)1.5–2cm, base persistent.

5 per fascicle, spreading to upcurved and ascending, persisting 5–6 years, 3–7cm × 1–1.5mm, pliant, dark green, abaxial surface with less conspicuous stomatal bands than adaxial surfaces, adaxial surfaces with strong, pale stomatal bands, margins finely serrulate, apex conic-acute to acuminate;

sheath 1–1.5(–2)cm, shed early.

Pollen cones

ellipsoid-cylindric, 10–15mm, orange-brown.

broadly ellipsoid-cylindric, ca. 15mm, pale red or yellow.

Seed(s)

cones maturing in 2 years, serotinous, long-persistent, remaining closed for 20 years or more, or opening on burning, in whorls, hard and heavy, very asymmetric, lanceoloid before opening, ovoid-cylindric when open, 8–15cm, yellow- or pale red-brown, stalks to 1cm;

apophyses toward outside base increasingly elongate, mammillate or raised-angled-conic, downcurved near base, scarcely raised on branchlet side, rhombic;

umbo central, low-pyramidal, sharp, upcurved.

cones maturing in 2 years, shedding seeds and falling soon thereafter, spreading, symmetric, lance-ovoid before opening, cylindro-ovoid when open, 7–15cm, straw-colored, resinous, sessile to short-stalked, apophyses much thickened, strongly cross-keeled, umbo terminal, depressed.

2n

=24.

=24.

Pinus attenuata

Pinus flexilis

Habitat Fire successional on dry slopes and foothills of Sierra Nevada and the Cascade and Coast ranges High montane forests, often at timberline
Elevation 300–1200m (1000–3900ft) (1000–)1500–3600m ((3300–)4900–11800ft)
Distribution
from FNA
CA; OR; Mexico in Baja California
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[BONAP county map]
from FNA
AZ; CA; CO; ID; MT; ND; NE; NM; NV; OR; SD; UT; WY; AB; BC
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Discussion

Pinus attenuata, mostly a chaparral species, bears cones at an early age. Its seed crops are heavy, and a hot fire permits the seeds to be released. It forms hybrids with P. muricata and P. radiata.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Pinus flexilis, much branched with a strongly tapering trunk, is little utilized because of its form and relative inaccessibility. It reportedly forms intermediates with P. strobiformis where the two overlap. The fresh-cut wood has the odor of turpentine.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 2. FNA vol. 2.
Parent taxa Pinaceae > Pinus Pinaceae > Pinus
Sibling taxa
P. albicaulis, P. aristata, P. balfouriana, P. banksiana, P. cembroides, P. clausa, P. contorta, P. coulteri, P. echinata, P. edulis, P. elliottii, P. engelmannii, P. flexilis, P. glabra, P. jeffreyi, P. lambertiana, P. leiophylla, P. longaeva, P. monophylla, P. monticola, P. muricata, P. palustris, P. ponderosa, P. pungens, P. quadrifolia, P. radiata, P. resinosa, P. rigida, P. sabiniana, P. serotina, P. strobiformis, P. strobus, P. sylvestris, P. taeda, P. torreyana, P. virginiana, P. washoensis
P. albicaulis, P. aristata, P. attenuata, P. balfouriana, P. banksiana, P. cembroides, P. clausa, P. contorta, P. coulteri, P. echinata, P. edulis, P. elliottii, P. engelmannii, P. glabra, P. jeffreyi, P. lambertiana, P. leiophylla, P. longaeva, P. monophylla, P. monticola, P. muricata, P. palustris, P. ponderosa, P. pungens, P. quadrifolia, P. radiata, P. resinosa, P. rigida, P. sabiniana, P. serotina, P. strobiformis, P. strobus, P. sylvestris, P. taeda, P. torreyana, P. virginiana, P. washoensis
Synonyms P. tuberculata Apinus flexilis
Name authority Lemmon: Mining Sci. Press 64: 45. (1892) E. James: Account Exped. Pittsburgh 2: 27, 35. (1823)
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