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knobcone pine

Apache pine, Arizona longleaf pine, pino real

Habit Shrubs or trees to 24m; trunk to 0.8m diam., usually straight; crown mostly narrowly to broadly conic. Trees to 35m; trunk to 0.6m diam., straight; crown irregularly rounded, rather thin.
Bark

purple-brown to dark brown, shallowly and narrowly fissured, with irregular, flat, loose-scaly plates, on upper sections of trunk nearly smooth.

dark brown, at maturity deeply furrowed, ridges becoming yellowish, of narrow, elongate, scaly plates.

Branches

ascending;

twigs slender, red-brown.

straight to ascending;

twigs stout (1–2cm thick), pale gray-brown, aging darker brown, rough.

Buds

ovoid to ovoid-cylindric, dark red-brown, aging darker, ca. 1.5cm, resinous;

scale margins fringed, apex attenuate.

ovoid-conic, to 2cm, resinous;

scale margins pale fringed.

Leaves

3 per fascicle, spreading or ascending, persisting 4–5 years, (8–)9–15(–20)cm × (1–)1.3–1.8mm, straight or slightly curved, twisted, yellow-green, all surfaces with fine stomatal lines, margins serrulate, apex abruptly conic-subulate;

sheath (1–)1.5–2cm, base persistent.

3(–5) per fascicle, spreading-ascending, often drooping, forming a brush at twig tips, persisting 2 years, (20–)25–45cm × 2mm, dull green, all surfaces with fine stomatal lines, margins coarsely serrulate, apex conic-subulate;

sheath 3–4cm, base persistent.

Pollen cones

ellipsoid-cylindric, 10–15mm, orange-brown.

cylindric, ca. 25mm, yellow to yellow-brown.

Seed(s)

cones maturing in 2 years, serotinous, long-persistent, remaining closed for 20 years or more, or opening on burning, in whorls, hard and heavy, very asymmetric, lanceoloid before opening, ovoid-cylindric when open, 8–15cm, yellow- or pale red-brown, stalks to 1cm;

apophyses toward outside base increasingly elongate, mammillate or raised-angled-conic, downcurved near base, scarcely raised on branchlet side, rhombic;

umbo central, low-pyramidal, sharp, upcurved.

cones maturing in 2 years and shedding seeds soon thereafter, not persistent, terminal, sometimes curved, often asymmetric, lance-ovoid before opening, ovoid when open, 11–14cm, light dull brown, nearly sessile or short-stalked;

apophyses rhombic, somewhat to quite elongate, strongly raised toward outer cone base, sometimes curved, strongly cross-keeled, narrowed to thick, curved, broadly triangular-based umbo, this often producing outcurved claw.

2n

=24.

=24.

Pinus attenuata

Pinus engelmannii

Habitat Fire successional on dry slopes and foothills of Sierra Nevada and the Cascade and Coast ranges High and dry mountain ranges, valleys, and plateaus
Elevation 300–1200m (1000–3900ft) 1500–2500m (4900–8200ft)
Distribution
from FNA
CA; OR; Mexico in Baja California
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[BONAP county map]
from FNA
AZ; NM; Mexico
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[BONAP county map]
Discussion

Pinus attenuata, mostly a chaparral species, bears cones at an early age. Its seed crops are heavy, and a hot fire permits the seeds to be released. It forms hybrids with P. muricata and P. radiata.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

In general appearance Pinus engelmannii much resembles P. palustris with its short-persistent, long leaves (but in this species drooping) and in its tendency to form a grass stage. It has a deep taproot as do P. palustris and P. ponderosa.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 2. FNA vol. 2.
Parent taxa Pinaceae > Pinus Pinaceae > Pinus
Sibling taxa
P. albicaulis, P. aristata, P. balfouriana, P. banksiana, P. cembroides, P. clausa, P. contorta, P. coulteri, P. echinata, P. edulis, P. elliottii, P. engelmannii, P. flexilis, P. glabra, P. jeffreyi, P. lambertiana, P. leiophylla, P. longaeva, P. monophylla, P. monticola, P. muricata, P. palustris, P. ponderosa, P. pungens, P. quadrifolia, P. radiata, P. resinosa, P. rigida, P. sabiniana, P. serotina, P. strobiformis, P. strobus, P. sylvestris, P. taeda, P. torreyana, P. virginiana, P. washoensis
P. albicaulis, P. aristata, P. attenuata, P. balfouriana, P. banksiana, P. cembroides, P. clausa, P. contorta, P. coulteri, P. echinata, P. edulis, P. elliottii, P. flexilis, P. glabra, P. jeffreyi, P. lambertiana, P. leiophylla, P. longaeva, P. monophylla, P. monticola, P. muricata, P. palustris, P. ponderosa, P. pungens, P. quadrifolia, P. radiata, P. resinosa, P. rigida, P. sabiniana, P. serotina, P. strobiformis, P. strobus, P. sylvestris, P. taeda, P. torreyana, P. virginiana, P. washoensis
Synonyms P. tuberculata P. macrophylla, P. apacheca, P. latifolia
Name authority Lemmon: Mining Sci. Press 64: 45. (1892) Carrière
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