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knobcone pine

slash pine

Habit Shrubs or trees to 24m; trunk to 0.8m diam., usually straight; crown mostly narrowly to broadly conic. Trees to 30m; trunk to 0.8m diam., straight to contorted; crown conic, becoming rounded or flattened.
Bark

purple-brown to dark brown, shallowly and narrowly fissured, with irregular, flat, loose-scaly plates, on upper sections of trunk nearly smooth.

orange- to purple-brown, irregularly furrowed and cross-checked into large, irregularly rectangular, papery-scaly plates.

Branches

ascending;

twigs slender, red-brown.

spreading to ascending;

twigs stout (to ca. 1cm thick), orange-brown, aging darker brown, rough-scaly.

Buds

ovoid to ovoid-cylindric, dark red-brown, aging darker, ca. 1.5cm, resinous;

scale margins fringed, apex attenuate.

cylindric, silvery brown, 1.5–2cm;

scale margins fringed.

Leaves

3 per fascicle, spreading or ascending, persisting 4–5 years, (8–)9–15(–20)cm × (1–)1.3–1.8mm, straight or slightly curved, twisted, yellow-green, all surfaces with fine stomatal lines, margins serrulate, apex abruptly conic-subulate;

sheath (1–)1.5–2cm, base persistent.

2 or 3 per fascicle, spreading or ascending, persisting ca. 2 years, 15–20(–23)cm × 1.2–1.5mm, straight, slightly twisted, pliant, yellow- to blue-green, all surfaces with stomatal lines, margins finely serrulate, apex abruptly acute to acuminate;

sheath 1–2cm, base persistent.

Pollen cones

ellipsoid-cylindric, 10–15mm, orange-brown.

cylindric, 30–40mm, purplish.

Seed(s)

cones maturing in 2 years, serotinous, long-persistent, remaining closed for 20 years or more, or opening on burning, in whorls, hard and heavy, very asymmetric, lanceoloid before opening, ovoid-cylindric when open, 8–15cm, yellow- or pale red-brown, stalks to 1cm;

apophyses toward outside base increasingly elongate, mammillate or raised-angled-conic, downcurved near base, scarcely raised on branchlet side, rhombic;

umbo central, low-pyramidal, sharp, upcurved.

cones maturing in 2 years, falling the year after seed-shed, single or in pairs, symmetric, lance-ovoid before opening, ovoid or ovoid-cylindric when open, (7–)9–18(–20)cm, light chocolate brown, on stalks to 3cm;

apophyses lustrous (as if varnished), slightly raised, strongly cross-keeled;

umbo central, depressed-pyramidal, with short, stout prickle.

2n

=24.

Pinus attenuata

Pinus elliottii

Habitat Fire successional on dry slopes and foothills of Sierra Nevada and the Cascade and Coast ranges
Elevation 300–1200m (1000–3900ft)
Distribution
from FNA
CA; OR; Mexico in Baja California
[WildflowerSearch map]
[BONAP county map]
from FNA
AL; FL; GA; LA; MS; SC [Introduced in subtropical and warm temperate areas worldwide]
[WildflowerSearch map]
[BONAP county map]
Discussion

Pinus attenuata, mostly a chaparral species, bears cones at an early age. Its seed crops are heavy, and a hot fire permits the seeds to be released. It forms hybrids with P. muricata and P. radiata.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Varieties 2 (native only in the flora).

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Key
1. Seedlings essentially without grass stage, buds thus scattered on the stem; leaves mostly in 3s, sometimes in 2s on same shoot; resin canals 3-5 per leaf; base of open cone ± truncate.
var. elliottii
1. Seedlings tending toward a grass stage, buds thus crowded on contracted stems; leaves mostly in 2s, sometimes in 3s on same shoot; resin canals 3-9 per leaf; base of open cone rounded.
var. densa
Source FNA vol. 2. FNA vol. 2.
Parent taxa Pinaceae > Pinus Pinaceae > Pinus
Sibling taxa
P. albicaulis, P. aristata, P. balfouriana, P. banksiana, P. cembroides, P. clausa, P. contorta, P. coulteri, P. echinata, P. edulis, P. elliottii, P. engelmannii, P. flexilis, P. glabra, P. jeffreyi, P. lambertiana, P. leiophylla, P. longaeva, P. monophylla, P. monticola, P. muricata, P. palustris, P. ponderosa, P. pungens, P. quadrifolia, P. radiata, P. resinosa, P. rigida, P. sabiniana, P. serotina, P. strobiformis, P. strobus, P. sylvestris, P. taeda, P. torreyana, P. virginiana, P. washoensis
P. albicaulis, P. aristata, P. attenuata, P. balfouriana, P. banksiana, P. cembroides, P. clausa, P. contorta, P. coulteri, P. echinata, P. edulis, P. engelmannii, P. flexilis, P. glabra, P. jeffreyi, P. lambertiana, P. leiophylla, P. longaeva, P. monophylla, P. monticola, P. muricata, P. palustris, P. ponderosa, P. pungens, P. quadrifolia, P. radiata, P. resinosa, P. rigida, P. sabiniana, P. serotina, P. strobiformis, P. strobus, P. sylvestris, P. taeda, P. torreyana, P. virginiana, P. washoensis
Subordinate taxa
P. elliottii var. densa, P. elliottii var. elliottii
Synonyms P. tuberculata P. heterophylla, P. taeda var. heterophylla
Name authority Lemmon: Mining Sci. Press 64: 45. (1892) Engelmann: Trans. Acad. Sci. St. Louis 4: 186, plates 1–3. (1880)
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