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knobcone pine

lodgepole pine, scrub pine, shore pine

Habit Shrubs or trees to 24m; trunk to 0.8m diam., usually straight; crown mostly narrowly to broadly conic. Shrubs or trees to 50m; trunk to 0.9m diam., straight to contorted; crown various according to genetic race.
Bark

purple-brown to dark brown, shallowly and narrowly fissured, with irregular, flat, loose-scaly plates, on upper sections of trunk nearly smooth.

brown to gray- or red-brown, platy to furrowed.

Branches

ascending;

twigs slender, red-brown.

Twigs

slender, orange to red-brown, aging darker brown, rough.

Buds

ovoid to ovoid-cylindric, dark red-brown, aging darker, ca. 1.5cm, resinous;

scale margins fringed, apex attenuate.

narrowly to broadly ovoid, dark red-brown, to 1.2cm, slightly resinous.

Leaves

3 per fascicle, spreading or ascending, persisting 4–5 years, (8–)9–15(–20)cm × (1–)1.3–1.8mm, straight or slightly curved, twisted, yellow-green, all surfaces with fine stomatal lines, margins serrulate, apex abruptly conic-subulate;

sheath (1–)1.5–2cm, base persistent.

2 per fascicle, spreading or ascending, persisting 3–8 years, 2–8cm × 0.7–2(–3)mm, twisted, yellow-green to dark green, all surfaces with fine stomatal lines, margins finely serrulate, apex blunt to acute or narrowly acuminate;

sheath 0.3–0.6(–1)cm, persistent.

Pollen cones

ellipsoid-cylindric, 10–15mm, orange-brown.

ellipsoid to cylindric, 5–15mm, orange-red.

Seed(s)

cones maturing in 2 years, serotinous, long-persistent, remaining closed for 20 years or more, or opening on burning, in whorls, hard and heavy, very asymmetric, lanceoloid before opening, ovoid-cylindric when open, 8–15cm, yellow- or pale red-brown, stalks to 1cm;

apophyses toward outside base increasingly elongate, mammillate or raised-angled-conic, downcurved near base, scarcely raised on branchlet side, rhombic;

umbo central, low-pyramidal, sharp, upcurved.

cones maturing in 2 years or variably serotinous, variably persistent, spreading to reflexed, often curved, nearly symmetric or variably asymmetric, lanceoloid to ovoid before opening, broadly ovoid to nearly globose when open, 2–6cm, tan to pale red-brown, lustrous, nearly sessile or on stalks to 1cm;

apophyses nearly rhombic, variously elongate, cross-keeled, often mammillate toward outer cone base and on inside above middle;

umbo central, depressed-triangular, prickle barely elongate to stubby or slender and to 6mm.

Lower

branches often descending, the upper spreading or ascending.

2n

=24.

=24 (variety not indicated).

Pinus attenuata

Pinus contorta

Habitat Fire successional on dry slopes and foothills of Sierra Nevada and the Cascade and Coast ranges
Elevation 300–1200m (1000–3900ft)
Distribution
from FNA
CA; OR; Mexico in Baja California
[WildflowerSearch map]
[BONAP county map]
from FNA
AK; CA; CO; ID; MT; NV; OR; SD; UT; WA; WY; AB; BC; NT; SK; YT; only in the flora
[WildflowerSearch map]
[BONAP county map]
Discussion

Pinus attenuata, mostly a chaparral species, bears cones at an early age. Its seed crops are heavy, and a hot fire permits the seeds to be released. It forms hybrids with P. muricata and P. radiata.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Pinus contorta is fire successional over most of its range and is characterized by prolific seeding and high seed viability in disturbed habitats, often resulting in extremely slow-growing, overly dense stands. Some authors consider it to consist of 4 races; these have been given various infraspecific ranks, but perhaps they are more conventionally treated as 3 varieties.

Varieties 3.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Key
1. Leaves 2-7 cm × 0.7-0.9(-1.1) mm, dark green; mature trunk with bark evidently furrowed; seed cones strongly asymmetric, strongly recurved, persistent or variously serotinous.
var. contorta
1. Leaves (4-)5-8 cm × (0.7-)1-2(-3) mm, yellow-green; mature trunk with bark not evidently furrowed; seed cones asymmetric to nearly symmetric, recurved to spreading, variously serotinous or soon shed.
→ 2
2. Seed cones asymmetric, recurved, variously serotinous, long-persistent; mid and lower apophyses mostly much domed; main branches mostly horizontally spreading, not ascending at tip.
var. latifolia
2. Seed cones nearly symmetric, mostly spreading, not serotinous, not persistent; mid and lower apophyses mostly shallowly domed; main branches ascending at tips.
var. murrayana
Source FNA vol. 2. FNA vol. 2.
Parent taxa Pinaceae > Pinus Pinaceae > Pinus
Sibling taxa
P. albicaulis, P. aristata, P. balfouriana, P. banksiana, P. cembroides, P. clausa, P. contorta, P. coulteri, P. echinata, P. edulis, P. elliottii, P. engelmannii, P. flexilis, P. glabra, P. jeffreyi, P. lambertiana, P. leiophylla, P. longaeva, P. monophylla, P. monticola, P. muricata, P. palustris, P. ponderosa, P. pungens, P. quadrifolia, P. radiata, P. resinosa, P. rigida, P. sabiniana, P. serotina, P. strobiformis, P. strobus, P. sylvestris, P. taeda, P. torreyana, P. virginiana, P. washoensis
P. albicaulis, P. aristata, P. attenuata, P. balfouriana, P. banksiana, P. cembroides, P. clausa, P. coulteri, P. echinata, P. edulis, P. elliottii, P. engelmannii, P. flexilis, P. glabra, P. jeffreyi, P. lambertiana, P. leiophylla, P. longaeva, P. monophylla, P. monticola, P. muricata, P. palustris, P. ponderosa, P. pungens, P. quadrifolia, P. radiata, P. resinosa, P. rigida, P. sabiniana, P. serotina, P. strobiformis, P. strobus, P. sylvestris, P. taeda, P. torreyana, P. virginiana, P. washoensis
Subordinate taxa
P. contorta var. contorta, P. contorta var. latifolia, P. contorta var. murrayana
Synonyms P. tuberculata
Name authority Lemmon: Mining Sci. Press 64: 45. (1892) Douglas ex Loudon: Arbor. Frutic. Brit. 4: 2292, figs. 2210, 2211. (1838)
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