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knobcone pine

Jack pine, pin gris, scrub pine

Habit Shrubs or trees to 24m; trunk to 0.8m diam., usually straight; crown mostly narrowly to broadly conic. Trees to 27m; trunk to 0.6m diam., straight to crooked; crown becoming irregularly rounded or spreading and flattened.
Bark

purple-brown to dark brown, shallowly and narrowly fissured, with irregular, flat, loose-scaly plates, on upper sections of trunk nearly smooth.

orange- to red-brown, scaly.

Branches

ascending;

twigs slender, red-brown.

descending to spreading-ascending, poorly self-pruning;

twigs slender, orange-red to red-brown, aging gray-brown, rough.

Buds

ovoid to ovoid-cylindric, dark red-brown, aging darker, ca. 1.5cm, resinous;

scale margins fringed, apex attenuate.

ovoid, red-brown, 0.5–1cm, resinous;

scale margins nearly entire.

Leaves

3 per fascicle, spreading or ascending, persisting 4–5 years, (8–)9–15(–20)cm × (1–)1.3–1.8mm, straight or slightly curved, twisted, yellow-green, all surfaces with fine stomatal lines, margins serrulate, apex abruptly conic-subulate;

sheath (1–)1.5–2cm, base persistent.

2 per fascicle, spreading or ascending, persisting 2–3 years, 2–5cm × 1–1.5(–2)mm, twisted, yellow-green, all surfaces with fine stomatal lines, margins finely serrulate, apex acute to short-subulate;

sheath 0.3–0.6cm, semipersistent.

Pollen cones

ellipsoid-cylindric, 10–15mm, orange-brown.

cylindric, 10–15mm, yellow to orange-brown.

Seed(s)

cones maturing in 2 years, serotinous, long-persistent, remaining closed for 20 years or more, or opening on burning, in whorls, hard and heavy, very asymmetric, lanceoloid before opening, ovoid-cylindric when open, 8–15cm, yellow- or pale red-brown, stalks to 1cm;

apophyses toward outside base increasingly elongate, mammillate or raised-angled-conic, downcurved near base, scarcely raised on branchlet side, rhombic;

umbo central, low-pyramidal, sharp, upcurved.

cones maturing in 2 years, shedding seeds soon thereafter or often long-serotinous and shedding seeds only through age or fire, upcurved, asymmetric, lanceoloid before opening, ovoid when open, 3–5.5cm, tan to light brown or greenish yellow, slick, nearly sessile or short-stalked, most apophyses depressed but increasingly mammillate toward outer cone base;

umbo central, depressed, small, sunken centrally, unarmed or with a small, reflexed apiculus.

2n

=24.

=24.

Pinus attenuata

Pinus banksiana

Habitat Fire successional on dry slopes and foothills of Sierra Nevada and the Cascade and Coast ranges Fire successional in boreal forests, tundra transition, dry flats, and hills, sandy soils
Elevation 300–1200m (1000–3900ft) 0–800m (0–2600ft)
Distribution
from FNA
CA; OR; Mexico in Baja California
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[BONAP county map]
from FNA
IL; IN; ME; MI; MN; NH; NY; PA; VT; WI; AB; BC; MB; NB; NS; NT; ON; PE; QC; SK
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[BONAP county map]
Discussion

Pinus attenuata, mostly a chaparral species, bears cones at an early age. Its seed crops are heavy, and a hot fire permits the seeds to be released. It forms hybrids with P. muricata and P. radiata.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Pinus banksiana reaches its largest size and best form in Canada. In western Alberta and in northeastern British Columbia, it is sympatric with P. contorta and forms hybrid swarms with that species.

Jack pine (Pinus banksiana) is the territorial tree of the Northwest Territories.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 2. FNA vol. 2.
Parent taxa Pinaceae > Pinus Pinaceae > Pinus
Sibling taxa
P. albicaulis, P. aristata, P. balfouriana, P. banksiana, P. cembroides, P. clausa, P. contorta, P. coulteri, P. echinata, P. edulis, P. elliottii, P. engelmannii, P. flexilis, P. glabra, P. jeffreyi, P. lambertiana, P. leiophylla, P. longaeva, P. monophylla, P. monticola, P. muricata, P. palustris, P. ponderosa, P. pungens, P. quadrifolia, P. radiata, P. resinosa, P. rigida, P. sabiniana, P. serotina, P. strobiformis, P. strobus, P. sylvestris, P. taeda, P. torreyana, P. virginiana, P. washoensis
P. albicaulis, P. aristata, P. attenuata, P. balfouriana, P. cembroides, P. clausa, P. contorta, P. coulteri, P. echinata, P. edulis, P. elliottii, P. engelmannii, P. flexilis, P. glabra, P. jeffreyi, P. lambertiana, P. leiophylla, P. longaeva, P. monophylla, P. monticola, P. muricata, P. palustris, P. ponderosa, P. pungens, P. quadrifolia, P. radiata, P. resinosa, P. rigida, P. sabiniana, P. serotina, P. strobiformis, P. strobus, P. sylvestris, P. taeda, P. torreyana, P. virginiana, P. washoensis
Synonyms P. tuberculata P. divaricata, P. sylvestris var. divaricata
Name authority Lemmon: Mining Sci. Press 64: 45. (1892) Lambert: Descr. Pinus 1: 7, plate 3. (1803)
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