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bristlecone pine, Colorado bristlecone pine, Rocky Mountain bristlecone pine

knobcone pine

Habit Trees to 15m; trunk to 1m diam., strongly tapering, twisted; crown rounded, flattened (sheared), or irregular. Shrubs or trees to 24m; trunk to 0.8m diam., usually straight; crown mostly narrowly to broadly conic.
Bark

gray to red-brown, shallowly fissured, with long, flat, irregular ridges.

purple-brown to dark brown, shallowly and narrowly fissured, with irregular, flat, loose-scaly plates, on upper sections of trunk nearly smooth.

Branches

contorted;

twigs pale red-brown, aging gray, puberulent, young branches resembling long bottlebrushes because of persistent leaves.

ascending;

twigs slender, red-brown.

Buds

ovoid-acuminate, pale red-brown, ca. 1cm, resinous.

ovoid to ovoid-cylindric, dark red-brown, aging darker, ca. 1.5cm, resinous;

scale margins fringed, apex attenuate.

Leaves

5 per fascicle, upcurved, persisting 10–17 years, (2–)3–4cm × 0.8–1mm, mostly connivent, deep blue-green, with drops and scales of resin, abaxial surface with strong, narrow median groove, adaxial surfaces conspicuously whitened by stomates, margins entire or distantly serrulate, apex conic-acute to conic-subulate;

sheath 0.5–1.5cm, scales soon recurving, shed early.

3 per fascicle, spreading or ascending, persisting 4–5 years, (8–)9–15(–20)cm × (1–)1.3–1.8mm, straight or slightly curved, twisted, yellow-green, all surfaces with fine stomatal lines, margins serrulate, apex abruptly conic-subulate;

sheath (1–)1.5–2cm, base persistent.

Pollen cones

ellipsoid, ca. 10mm, bluish to red.

ellipsoid-cylindric, 10–15mm, orange-brown.

Seed(s)

cones maturing in 2 years, shedding seeds and falling soon thereafter, spreading, symmetric, lance-cylindric before opening, lance-ovoid to ovoid or cylindric when open, 6–11cm, purple to brown, nearly sessile;

apophyses much thickened;

umbo central, with triangular base, extended into slender, brittle prickle 4–10mm.

cones maturing in 2 years, serotinous, long-persistent, remaining closed for 20 years or more, or opening on burning, in whorls, hard and heavy, very asymmetric, lanceoloid before opening, ovoid-cylindric when open, 8–15cm, yellow- or pale red-brown, stalks to 1cm;

apophyses toward outside base increasingly elongate, mammillate or raised-angled-conic, downcurved near base, scarcely raised on branchlet side, rhombic;

umbo central, low-pyramidal, sharp, upcurved.

2n

=24.

=24.

Pinus aristata

Pinus attenuata

Habitat Subalpine and alpine Fire successional on dry slopes and foothills of Sierra Nevada and the Cascade and Coast ranges
Elevation 2500–3400m (8200–11200ft) 300–1200m (1000–3900ft)
Distribution
from FNA
AZ; CO; NM
[WildflowerSearch map]
[BONAP county map]
from FNA
CA; OR; Mexico in Baja California
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[BONAP county map]
Discussion

Pinus aristata has leaves usually narrower and sharper than in P. longaeva and P. balfouriana, and the leaves almost always have a narrow, median groove on the abaxial surface.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Pinus attenuata, mostly a chaparral species, bears cones at an early age. Its seed crops are heavy, and a hot fire permits the seeds to be released. It forms hybrids with P. muricata and P. radiata.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 2. FNA vol. 2.
Parent taxa Pinaceae > Pinus Pinaceae > Pinus
Sibling taxa
P. albicaulis, P. attenuata, P. balfouriana, P. banksiana, P. cembroides, P. clausa, P. contorta, P. coulteri, P. echinata, P. edulis, P. elliottii, P. engelmannii, P. flexilis, P. glabra, P. jeffreyi, P. lambertiana, P. leiophylla, P. longaeva, P. monophylla, P. monticola, P. muricata, P. palustris, P. ponderosa, P. pungens, P. quadrifolia, P. radiata, P. resinosa, P. rigida, P. sabiniana, P. serotina, P. strobiformis, P. strobus, P. sylvestris, P. taeda, P. torreyana, P. virginiana, P. washoensis
P. albicaulis, P. aristata, P. balfouriana, P. banksiana, P. cembroides, P. clausa, P. contorta, P. coulteri, P. echinata, P. edulis, P. elliottii, P. engelmannii, P. flexilis, P. glabra, P. jeffreyi, P. lambertiana, P. leiophylla, P. longaeva, P. monophylla, P. monticola, P. muricata, P. palustris, P. ponderosa, P. pungens, P. quadrifolia, P. radiata, P. resinosa, P. rigida, P. sabiniana, P. serotina, P. strobiformis, P. strobus, P. sylvestris, P. taeda, P. torreyana, P. virginiana, P. washoensis
Synonyms P. balfouriana var. aristata P. tuberculata
Name authority Engelmann: in Parry & Engelmann, Amer. J. Sci. Arts ser. 2, 34: 331. (1862) Lemmon: Mining Sci. Press 64: 45. (1892)
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