Pieris |
Ericaceae subfam. vaccinioideae |
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fetterbush |
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Habit | Shrubs [trees] or vines. | |||||
Stems | ± erect; twigs multicellular stipitate-glandular-hairy and, sometimes, conspicuously strigose (hairs elongate, stiff, long-celled, eglandular), otherwise sparsely to densely unicellular-hairy. |
erect, arching, spreading, creeping, or procumbent. |
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Leaves | persistent or deciduous, alternate, sometimes pseudoverticillate [in whorls of 3]; blade ovate, elliptic, or slightly obovate, coriaceous, margins obscurely to clearly toothed or serrulate [entire], plane or revolute, surfaces multicellular, short-stalked stipitate-glandular-hairy (along with, in P. floribunda, stout, elongate hairs associated with marginal teeth, and unicellular-hairy on midvein adaxially); venation brochidodromous to reticulodromous (veins of varied thickness and conspicuousness). |
deciduous or persistent, alternate, sometimes pseudoverticillate (Pieris); petiole usually present, sometimes absent (some species of Vaccinium); blade plane, abaxial groove absent. |
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Inflorescences | axillary or terminal panicles or racemes, 15–35-flowered, (borne the year preceding flowering). |
usually axillary, sometimes terminal, usually panicles or racemes, sometimes corymbs or fascicles, sometimes solitary flowers, (borne on leafy twigs, except Zenobia on leafless twigs); perulae absent; bracts much shorter than sepals (sometimes absent). |
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Pedicels | bracteoles 2, proximal or distal. |
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Flowers | sepals 5, slightly connate, ovate [oblong-lanceolate]; petals 5, connate for nearly their entire lengths, white, corolla urceolate to cylindric-urceolate, lobes short; stamens 10, included; filaments straight or geniculate, glabrous or hairy, with 2 stout, minutely papillose spurs at anther-filament junction; anthers without awns, dehiscent by elliptic pores; pistil 5-carpellate; ovary 5-locular; stigma capitate. |
pendulous; perianth and androecium hypogynous or epigynous (Gaylussacia, Vaccinium); sepals (4-)5[-8]; petals 4-5(-6), connate (rarely distinct or nearly so in some species of Vaccinium), corolla deciduous, campanulate, cylindric, or urceolate, lobes usually much shorter (sometimes longer) than tube; intrastaminal nectary disc absent or present; stamens 8-10[-16]; anthers dehiscent by terminal pores or short slits; ovary 5- or 10-locular; placentation axile; style straight. |
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Fruits | capsular, subglobose or globose to ovoid, (with unthickened sutures), dry. |
capsular, dehiscence loculicidal, or baccate or drupaceous, indehiscent. |
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Seeds | 40–100, ellipsoidal to obovoid, ± rectangular or angular-ovoid, or narrowly conic, flattened or not, (sometimes slightly winged); testa cells elongate or isodiametric. |
2-300, distinct, ovoid or obovoid to ellipsoid, lanceoloid, or conic, to angular or wedge- or crescent-shaped, usually not winged, sometimes slightly winged or tailed. |
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x | = 12. |
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Pieris |
Ericaceae subfam. vaccinioideae |
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Distribution |
se United States; West Indies (Cuba); e Asia |
Nearly worldwide; especially arctic; temperate; and alpine areas; also very diverse in neotropical cloud forests |
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Discussion | Ampelothamnus Small; Arcterica Coville; Portuna Nuttall Species 7 (2 in the flora). (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Genera 46, species ca. 1600 (12 genera, 58 species in the flora) (Discussion copyrighted by Flora of North America; reprinted with permission.) |
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Key |
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Source | FNA vol. 8, p. 497. | FNA vol. 8, p. 496. | ||||
Parent taxa | ||||||
Subordinate taxa | ||||||
Name authority | D. Don: Edinburgh New Philos. J. 17: 159. 1834 , | Arnott: M. Napier, Encycl. Brit. ed. 7 5: 118. (1832) — (as Vaccinieae) | ||||
Web links |