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fetterbush

Habit Shrubs [trees] or vines.
Stems

± erect;

twigs multicellular stipitate-glandular-hairy and, sometimes, conspicuously strigose (hairs elongate, stiff, long-celled, eglandular), otherwise sparsely to densely unicellular-hairy.

erect, arching, spreading, creeping, or procumbent.

Leaves

persistent or deciduous, alternate, sometimes pseudoverticillate [in whorls of 3];

blade ovate, elliptic, or slightly obovate, coriaceous, margins obscurely to clearly toothed or serrulate [entire], plane or revolute, surfaces multicellular, short-stalked stipitate-glandular-hairy (along with, in P. floribunda, stout, elongate hairs associated with marginal teeth, and unicellular-hairy on midvein adaxially);

venation brochidodromous to reticulodromous (veins of varied thickness and conspicuousness).

deciduous or persistent, alternate, sometimes pseudoverticillate (Pieris);

petiole usually present, sometimes absent (some species of Vaccinium);

blade plane, abaxial groove absent.

Inflorescences

axillary or terminal panicles or racemes, 15–35-flowered, (borne the year preceding flowering).

usually axillary, sometimes terminal, usually panicles or racemes, sometimes corymbs or fascicles, sometimes solitary flowers, (borne on leafy twigs, except Zenobia on leafless twigs);

perulae absent;

bracts much shorter than sepals (sometimes absent).

Pedicels

bracteoles 2, proximal or distal.

Flowers

sepals 5, slightly connate, ovate [oblong-lanceolate];

petals 5, connate for nearly their entire lengths, white, corolla urceolate to cylindric-urceolate, lobes short;

stamens 10, included;

filaments straight or geniculate, glabrous or hairy, with 2 stout, minutely papillose spurs at anther-filament junction;

anthers without awns, dehiscent by elliptic pores;

pistil 5-carpellate;

ovary 5-locular;

stigma capitate.

pendulous;

perianth and androecium hypogynous or epigynous (Gaylussacia, Vaccinium);

sepals (4-)5[-8];

petals 4-5(-6), connate (rarely distinct or nearly so in some species of Vaccinium), corolla deciduous, campanulate, cylindric, or urceolate, lobes usually much shorter (sometimes longer) than tube;

intrastaminal nectary disc absent or present;

stamens 8-10[-16];

anthers dehiscent by terminal pores or short slits;

ovary 5- or 10-locular;

placentation axile;

style straight.

Fruits

capsular, subglobose or globose to ovoid, (with unthickened sutures), dry.

capsular, dehiscence loculicidal, or baccate or drupaceous, indehiscent.

Seeds

40–100, ellipsoidal to obovoid, ± rectangular or angular-ovoid, or narrowly conic, flattened or not, (sometimes slightly winged);

testa cells elongate or isodiametric.

2-300, distinct, ovoid or obovoid to ellipsoid, lanceoloid, or conic, to angular or wedge- or crescent-shaped, usually not winged, sometimes slightly winged or tailed.

x

= 12.

Pieris

Ericaceae subfam. vaccinioideae

Distribution
from USDA
se United States; West Indies (Cuba); e Asia
[BONAP county map]
Nearly worldwide; especially arctic; temperate; and alpine areas; also very diverse in neotropical cloud forests
Discussion

Ampelothamnus Small; Arcterica Coville; Portuna Nuttall

Species 7 (2 in the flora).

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Genera 46, species ca. 1600 (12 genera, 58 species in the flora)

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Key
1. Inflorescences axillary racemes; capsules with placentae central to nearly basal; styles strongly sunken into ovary apex; seeds with isodiametric testa cells; filaments geniculate, glabrous; twigs multicellular stipitate-glandular-hairy.
P. phillyreifolia
1. Inflorescences terminal panicles; capsules with placentae subapical; styles slightly sunken into ovary apex; seeds with elongated testa cells; filaments straight, hairy; twigs multicellular stipitate-glandular-hairy and conspicuously strigose (hairs stout, elongate).
P. floribunda
Source FNA vol. 8, p. 497. Author: Walter S. Judd. FNA vol. 8, p. 496. Author: Gordon C. Tucker.
Parent taxa Ericaceae > subfam. Vaccinioideae Ericaceae
Subordinate taxa
P. floribunda, P. phillyreifolia
Name authority D. Don: Edinburgh New Philos. J. 17: 159. 1834 , Arnott: M. Napier, Encycl. Brit. ed. 7 5: 118. (1832) — (as Vaccinieae)
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