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ninebark, Pacific nine-bark, tall ninebark

dwarf nine-bark, Nevada nine-bark

Habit Shrubs, to 45(–60) dm. Shrubs, 15 dm, densely branched.
Stems

erect, sometimes suckering, angled, glabrous or finely stellate-hairy.

orientation not recorded, brown becoming gray, stellate-hairy, glabrescent.

Leaves

stipules linear to narrowly elliptic, 4 × 0.5–2 mm;

petiole 1–2(–3) cm;

blade broadly ovate to obovate, (3–)4–8 cm, usually as wide as long, base rounded to truncate or slightly cordate, 3- or 5-lobed, margins irregularly crenate to doubly serrate, apex obtuse to acute, abaxial surface paler, glabrous or more densely stellate-hairy, adaxial glabrous or sparsely stellate-hairy.

stipules oblong to narrowly elliptic or rhombic, 2–4 × 1.5–2 mm;

petiole 2–10 mm;

blade broadly ovate to orbiculate, 0.5–2 × 0.5–2 cm, slightly longer than wide, base rounded to truncate or slightly cordate, (0)3- or 5(7)-lobed, margins crenate to doubly crenate, apex rounded, abaxial surface more densely stellate-hairy (especially on veins), adaxial sparsely so.

Inflorescences

30–50-flowered, dense, hemispheric racemes, 3 cm diam., sometimes compound with some proximal pedicels becoming secondary peduncles;

bracts narrowly elliptic to spatulate, 4 × 2 mm, apex acute or erose-dentate, faces glandular.

5–10(–15)-flowered, open, umbel-like racemes, 3 cm diam.;

bracts similar to stipules.

Pedicels

1–1.5 cm, densely stellate-hairy.

0.6–1.5 cm, densely stellate-hairy.

Flowers

5–8 mm diam.;

hypanthium cup-shaped, 2 mm, densely stellate-hairy;

sepals pale green to white, darker in center, triangular, 2–3 mm, apex gland-tipped, surfaces densely stellate-hairy;

petals white, broadly elliptic to orbiculate, 3–4 × 3–4 mm;

stamens equal to or exceeding petals;

carpels 3–5, connate basally, mostly glabrous, sometimes hairy (on ventral suture).

7 mm diam.;

hypanthium cup-shaped to campanulate, 1.5–3 mm, stellate-hairy;

sepals pale green to white, darker in center, triangular to narrowly ovate, 2.5–3 mm, apex gland-tipped, surfaces stellate-hairy;

petals white to pale pink, broadly obovate to orbiculate, 3 × 2.5 mm;

stamens 20–30, unequal (alternately long and short), shorter than petals, anthers pink to dark brown;

carpels 1(2), if 2 then connate at least 1/2 their lengths, indumentum not recorded.

Seeds

2(–5), pyriform, 2.3–2.8 mm.

1 or 2, 2 mm.

Follicles

3–5, connate basally, shiny brown, ovoid, 5–7 mm (lengths slightly exceeding sepals), glabrous;

styles 2.5–3 mm.

1(2), pale buff, shape not recorded, 3–6 mm (lengths equal to or slightly exceeding sepals), densely stellate-hairy;

styles 2.5 mm sometimes distinctly bent.

2n

= 18.

Physocarpus capitatus

Physocarpus alternans

Phenology Flowering May–Jul; fruiting Jul–Aug. Flowering Jun–Jul; fruiting Aug.
Habitat Open sunny slopes on clay (higher elevations), stream and swamp banks, lake margins in moist woods (lower elevations) Dry rocky canyon slopes, among pinyon pine, juniper, and Cercocarpus woodlands
Elevation 20–1300 m (100–4300 ft) 1500–3100 m (4900–10200 ft)
Distribution
from FNA
AK; CA; ID; OR; WA; BC
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[BONAP county map]
from FNA
CA; CO; ID; NV; UT; WY
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[BONAP county map]
Discussion

Physocarpus capitatus is commonly cultivated.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

The two subspecies described from California by Howell appear to be local variants with relatively small differences in leaf and hypanthium indumentum that probably do not merit formal recognition. Being drought tolerant, Physocarpus alternans should be considered more often for planting in dry habitats.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 9, p. 349. FNA vol. 9, p. 350.
Parent taxa Rosaceae > subfam. Amygdaloideae > tribe Neillieae > Physocarpus Rosaceae > subfam. Amygdaloideae > tribe Neillieae > Physocarpus
Sibling taxa
P. alternans, P. intermedius, P. malvaceus, P. monogynus, P. opulifolius
P. capitatus, P. intermedius, P. malvaceus, P. monogynus, P. opulifolius
Synonyms Spiraea capitata, P. opulifolius var. tomentellus Neillia monogyna var. alternans, P. alternans subsp. annulatus, P. alternans subsp. panamintensis
Name authority (Pursh) Kuntze: Revis. Gen. Pl. 1: 219. (1891) (M. E. Jones) J. T. Howell: Proc. Calif. Acad. Sci., ser. 4, 20: 130. (1931)
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