Physocarpus capitatus |
Physocarpus |
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ninebark, Pacific nine-bark, tall ninebark |
nine-bark, physocarpe |
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Habit | Shrubs, to 45(–60) dm. | Shrubs, (10–)30–60 dm, glabrous or stellate-hairy, mostly eglandular (except in inflorescence). | ||||||||||||||||||||
Stems | erect, sometimes suckering, angled, glabrous or finely stellate-hairy. |
5–20, ascending to erect, branches erect proximally, somewhat sinuous; bark buff to reddish brown or black, usually peeling in narrow strips; short shoots present; buds superposed, each with ca. 5 exposed brown scales. |
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Leaves | stipules linear to narrowly elliptic, 4 × 0.5–2 mm; petiole 1–2(–3) cm; blade broadly ovate to obovate, (3–)4–8 cm, usually as wide as long, base rounded to truncate or slightly cordate, 3- or 5-lobed, margins irregularly crenate to doubly serrate, apex obtuse to acute, abaxial surface paler, glabrous or more densely stellate-hairy, adaxial glabrous or sparsely stellate-hairy. |
deciduous, cauline; stipules caducous, decurrent on stem, leaving oblique, broadly ovate to rectangular stubs, linear or oblong to narrowly ovate, obovate, or elliptic, sometimes rhombic, margins sometimes remotely glandular-dentate, surfaces sparsely stellate-hairy (glabrous adaxially in P. monogynus); petiole present; blade broadly ovate to obovate or orbiculate, 0.5–10 cm, membranous, margins flat, crenate to doubly crenate or doubly dentate, 3–5(–7)-lobed, rarely almost unlobed, 3–5(–7)-veined, surfaces glabrous or sparsely to densely stellate-hairy. |
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Inflorescences | 30–50-flowered, dense, hemispheric racemes, 3 cm diam., sometimes compound with some proximal pedicels becoming secondary peduncles; bracts narrowly elliptic to spatulate, 4 × 2 mm, apex acute or erose-dentate, faces glandular. |
terminal or axillary on leafy short shoots, 5–50-flowered, open to dense corymb- or umbel-like pedunculate racemes; bracts present; bracteoles absent. |
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Pedicels | 1–1.5 cm, densely stellate-hairy. |
present. |
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Flowers | 5–8 mm diam.; hypanthium cup-shaped, 2 mm, densely stellate-hairy; sepals pale green to white, darker in center, triangular, 2–3 mm, apex gland-tipped, surfaces densely stellate-hairy; petals white, broadly elliptic to orbiculate, 3–4 × 3–4 mm; stamens equal to or exceeding petals; carpels 3–5, connate basally, mostly glabrous, sometimes hairy (on ventral suture). |
5–12 mm diam.; hypanthium cup-shaped to campanulate, 1.5–3 mm, exterior stellate-hairy; sepals 5, ascending to slightly reflexed, triangular to ovate; petals 5, white to pale pink, broadly obovate, elliptic, or orbiculate, base short-clawed, sometimes irregularly toothed or notched; stamens 20–40, shorter than, equal to, or exceeding petals; carpels 1–5; styles slender, stigmas capitate; ovules 2–5. |
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Fruits | aggregated follicles, sometimes solitary, 1–5, ovoid, often ± inflated, slightly flattened, size not recorded, splitting along both sutures; hypanthium persistent; sepals persistent, ascending to slightly reflexed. |
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Seeds | 2(–5), pyriform, 2.3–2.8 mm. |
1(–5), shiny yellow, carunculate. |
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Follicles | 3–5, connate basally, shiny brown, ovoid, 5–7 mm (lengths slightly exceeding sepals), glabrous; styles 2.5–3 mm. |
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x | = 9. |
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2n | = 18. |
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Physocarpus capitatus |
Physocarpus |
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Phenology | Flowering May–Jul; fruiting Jul–Aug. | |||||||||||||||||||||
Habitat | Open sunny slopes on clay (higher elevations), stream and swamp banks, lake margins in moist woods (lower elevations) | |||||||||||||||||||||
Elevation | 20–1300 m (100–4300 ft) | |||||||||||||||||||||
Distribution |
AK; CA; ID; OR; WA; BC
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North America; n Mexico; ne Asia [Introduced in Europe] |
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Discussion | Physocarpus capitatus is commonly cultivated. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Species 8–10 (6 in the flora). Some species of Physocarpus are not well delimited. Some botanists have confused Physocarpus and Neillia (including Stephanandra); Physocarpus is distinguished by its rounded umbel-like racemes and bladderlike fruits splitting along both sutures; in Neillia, the inflorescences are longer and narrower, and the fruits split along the adaxial suture. As currently understood, Neillia is an Asian genus. Two or three species of Physocarpus are widely cultivated in North America and Europe; the others are less commonly seen though they may be worthy of more attention. Opulaster Medikus ex Kuntze is an illegitimate, superfluous name that pertains here. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
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Key |
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Source | FNA vol. 9, p. 349. | FNA vol. 9, p. 347. | ||||||||||||||||||||
Parent taxa | Rosaceae > subfam. Amygdaloideae > tribe Neillieae > Physocarpus | Rosaceae > subfam. Amygdaloideae > tribe Neillieae | ||||||||||||||||||||
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Subordinate taxa | ||||||||||||||||||||||
Synonyms | Spiraea capitata, P. opulifolius var. tomentellus | Spiraea section P. | ||||||||||||||||||||
Name authority | (Pursh) Kuntze: Revis. Gen. Pl. 1: 219. (1891) | (Cambessèdes) Rafinesque: New Fl. 3: 73. (1838) | ||||||||||||||||||||
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