Physocarpus capitatus |
Rosaceae tribe Neillieae |
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ninebark, Pacific nine-bark, tall ninebark |
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Habit | Shrubs, to 45(–60) dm. | Shrubs; unarmed. | ||||
Stems | erect, sometimes suckering, angled, glabrous or finely stellate-hairy. |
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Leaves | stipules linear to narrowly elliptic, 4 × 0.5–2 mm; petiole 1–2(–3) cm; blade broadly ovate to obovate, (3–)4–8 cm, usually as wide as long, base rounded to truncate or slightly cordate, 3- or 5-lobed, margins irregularly crenate to doubly serrate, apex obtuse to acute, abaxial surface paler, glabrous or more densely stellate-hairy, adaxial glabrous or sparsely stellate-hairy. |
alternate, simple; stipules persistent or caducous, free; venation palmate. |
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Inflorescences | 30–50-flowered, dense, hemispheric racemes, 3 cm diam., sometimes compound with some proximal pedicels becoming secondary peduncles; bracts narrowly elliptic to spatulate, 4 × 2 mm, apex acute or erose-dentate, faces glandular. |
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Pedicels | 1–1.5 cm, densely stellate-hairy. |
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Flowers | 5–8 mm diam.; hypanthium cup-shaped, 2 mm, densely stellate-hairy; sepals pale green to white, darker in center, triangular, 2–3 mm, apex gland-tipped, surfaces densely stellate-hairy; petals white, broadly elliptic to orbiculate, 3–4 × 3–4 mm; stamens equal to or exceeding petals; carpels 3–5, connate basally, mostly glabrous, sometimes hairy (on ventral suture). |
perianth and androecium perigynous; epicalyx bractlets absent; hypanthium cup-shaped to campanulate; torus absent; carpels 1–5, basally connate (distinct when single), free, styles terminal, distinct; ovules 2–5[–10], marginal, pendulous, biseriate or clustered. |
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Fruits | solitary or aggregated follicles; styles persistent, not elongate. |
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Seeds | 2(–5), pyriform, 2.3–2.8 mm. |
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Follicles | 3–5, connate basally, shiny brown, ovoid, 5–7 mm (lengths slightly exceeding sepals), glabrous; styles 2.5–3 mm. |
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2n | = 18. |
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Physocarpus capitatus |
Rosaceae tribe Neillieae |
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Phenology | Flowering May–Jul; fruiting Jul–Aug. | |||||
Habitat | Open sunny slopes on clay (higher elevations), stream and swamp banks, lake margins in moist woods (lower elevations) | |||||
Elevation | 20–1300 m (100–4300 ft) | |||||
Distribution |
AK; CA; ID; OR; WA; BC
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North America; n Mexico; Asia [Introduced in Europe] |
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Discussion | Physocarpus capitatus is commonly cultivated. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Genera 2, species 21 (2 genera, 7 species in the flora). The base chromosome number for Neillieae is x = 9. Cyanogenic glycosides are absent in Physocarpus; there are no data on Neillia. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
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Key |
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Source | FNA vol. 9, p. 349. | FNA vol. 9, p. 347. | ||||
Parent taxa | Rosaceae > subfam. Amygdaloideae > tribe Neillieae > Physocarpus | Rosaceae > subfam. Amygdaloideae | ||||
Sibling taxa | ||||||
Subordinate taxa | ||||||
Synonyms | Spiraea capitata, P. opulifolius var. tomentellus | |||||
Name authority | (Pursh) Kuntze: Revis. Gen. Pl. 1: 219. (1891) | Maximowicz: Trudy Imp. S.-Peterburgsk. Bot. Sada 6: 164, 216. (1879) | ||||
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