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ninebark, Pacific nine-bark, tall ninebark

Habit Shrubs, to 45(–60) dm. Shrubs; unarmed.
Stems

erect, sometimes suckering, angled, glabrous or finely stellate-hairy.

Leaves

stipules linear to narrowly elliptic, 4 × 0.5–2 mm;

petiole 1–2(–3) cm;

blade broadly ovate to obovate, (3–)4–8 cm, usually as wide as long, base rounded to truncate or slightly cordate, 3- or 5-lobed, margins irregularly crenate to doubly serrate, apex obtuse to acute, abaxial surface paler, glabrous or more densely stellate-hairy, adaxial glabrous or sparsely stellate-hairy.

alternate, simple;

stipules persistent or caducous, free;

venation palmate.

Inflorescences

30–50-flowered, dense, hemispheric racemes, 3 cm diam., sometimes compound with some proximal pedicels becoming secondary peduncles;

bracts narrowly elliptic to spatulate, 4 × 2 mm, apex acute or erose-dentate, faces glandular.

Pedicels

1–1.5 cm, densely stellate-hairy.

Flowers

5–8 mm diam.;

hypanthium cup-shaped, 2 mm, densely stellate-hairy;

sepals pale green to white, darker in center, triangular, 2–3 mm, apex gland-tipped, surfaces densely stellate-hairy;

petals white, broadly elliptic to orbiculate, 3–4 × 3–4 mm;

stamens equal to or exceeding petals;

carpels 3–5, connate basally, mostly glabrous, sometimes hairy (on ventral suture).

perianth and androecium perigynous;

epicalyx bractlets absent;

hypanthium cup-shaped to campanulate;

torus absent;

carpels 1–5, basally connate (distinct when single), free, styles terminal, distinct;

ovules 2–5[–10], marginal, pendulous, biseriate or clustered.

Fruits

solitary or aggregated follicles;

styles persistent, not elongate.

Seeds

2(–5), pyriform, 2.3–2.8 mm.

Follicles

3–5, connate basally, shiny brown, ovoid, 5–7 mm (lengths slightly exceeding sepals), glabrous;

styles 2.5–3 mm.

2n

= 18.

Physocarpus capitatus

Rosaceae tribe Neillieae

Phenology Flowering May–Jul; fruiting Jul–Aug.
Habitat Open sunny slopes on clay (higher elevations), stream and swamp banks, lake margins in moist woods (lower elevations)
Elevation 20–1300 m (100–4300 ft)
Distribution
from FNA
AK; CA; ID; OR; WA; BC
[WildflowerSearch map]
[BONAP county map]
North America; n Mexico; Asia [Introduced in Europe]
Discussion

Physocarpus capitatus is commonly cultivated.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Genera 2, species 21 (2 genera, 7 species in the flora).

The base chromosome number for Neillieae is x = 9. Cyanogenic glycosides are absent in Physocarpus; there are no data on Neillia.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Key
1. Inflorescences corymb- or umbel-like racemes; carpels 1–5; fruits usually aggregated follicles.
Physocarpus
1. Inflorescences racemes or panicles to corymbs; carpel 1; fruits solitary follicles.
Neillia
Source FNA vol. 9, p. 349. FNA vol. 9, p. 347. Author: Luc Brouillet.
Parent taxa Rosaceae > subfam. Amygdaloideae > tribe Neillieae > Physocarpus Rosaceae > subfam. Amygdaloideae
Sibling taxa
P. alternans, P. intermedius, P. malvaceus, P. monogynus, P. opulifolius
Subordinate taxa
Neillia, Physocarpus
Synonyms Spiraea capitata, P. opulifolius var. tomentellus
Name authority (Pursh) Kuntze: Revis. Gen. Pl. 1: 219. (1891) Maximowicz: Trudy Imp. S.-Peterburgsk. Bot. Sada 6: 164, 216. (1879)
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