Physaria vitulifera |
Physaria tumulosa |
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roundtip twinpod |
Kodachrome bladderpod |
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Habit | Perennials; caudex simple or branched, (cespitose); (silvery) pubescent throughout, trichomes several-rayed, rays furcate or bifurcate, (relatively massive, smooth to few-tubercled). | Perennials; caudex (buried), branched, (forming hard mats); densely pubescent, trichomes several-rayed, rays furcate or bifurcate, (tuberculate). |
Stems | several from base, usually decumbent to ascending, (arising laterally, unbranched, coarse), 1–2 dm. |
several from base, erect, (unbranched), 0.2–0.3 dm. |
Basal leaves | blade pandurate or obovate, 3–6 cm, margins usually deeply and broadly incised, rarely subentire, (apex obtuse). |
(few), similar to cauline. |
Cauline leaves | blade oblanceolate to spatulate, similar to basal, (3–6 mm wide), margins entire, (apex often somewhat acute). |
(petiole not differentiated from blade); blade (somewhat succulent), linear to narrowly oblanceolate, 5–12 mm, margins entire. |
Racemes | congested, (elongated in fruit). |
dense, (few-flowered). |
Flowers | sepals oblong, 6–8 mm; petals spatulate, to 10 mm. |
sepals (yellowish), elliptic, 3–4.5 mm; petals (erect or, more commonly, arching), spatulate to oblanceolate, 5.8–7 mm, (claw not or weakly differentiated from blade). |
Fruiting pedicels | (usually curving upward, sigmoid), 6–10 mm. |
(ascending to divaricate-ascending, ± straight), 3.5–6 mm. |
Fruits | didymous, irregular in shape, somewhat angular, inflated, 5–7 × 6–8 mm, (papery, often rigid, base obtuse or truncate, apical sinus broad, open and deep); valves (retaining seeds after dehiscence), pubescent, trichomes spreading, loose; replum oblong, often constricted, as wide as or wider than fruit, apex obtuse; ovules 4 per ovary; style 5–7 mm. |
(coppery or reddish brown in age), broadly ovoid, slightly inflated, 3–4 mm; valves (not retaining seeds after dehiscence), glabrous throughout; replum as wide as or wider than fruit; ovules 4–8 per ovary; style 1.8–3 mm. |
Seeds | flattened. |
flattened. |
2n | = 8, 16. |
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Physaria vitulifera |
Physaria tumulosa |
|
Phenology | Flowering Apr–Jun. | Flowering May–Jun. |
Habitat | Rocky hillsides, dry banks, gravel and sand, granitic slopes, soil scree, red shale | Barren white knolls surrounded by sagebrush, pinyon pine, and Utah juniper |
Elevation | 1600-3000 m (5200-9800 ft) | 1600-1800 m (5200-5900 ft) |
Distribution |
CO
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UT |
Discussion | Of conservation concern. Physaria tumulosa is morphologically similar to 55. P. navajoensis of northeastern Arizona and northwestern New Mexico, and differing very subtly. It has been long treated as an infraspecific taxon of P. hitchcockii; unpublished molecular data do not support that disposition. It is found on knolls of the Winsor Member of the Carmel Formation. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
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Source | FNA vol. 7, p. 665. | FNA vol. 7, p. 664. |
Parent taxa | Brassicaceae > tribe Physarieae > Physaria | Brassicaceae > tribe Physarieae > Physaria |
Sibling taxa | ||
Synonyms | Lesquerella hitchcockii subsp. tumulosa, Lesquerella tumulosa, P. rubicundula var. tumulosa | |
Name authority | Rydberg: Bull. Torrey Bot. Club 28: 278. (1901) | (Barneby) O’Kane & Al-Shehbaz: Novon 12: 328. (2002) |
Web links |