Physaria vitulifera |
Physaria hitchcockii |
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roundtip twinpod |
Hitchcock's bladderpod |
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Habit | Perennials; caudex simple or branched, (cespitose); (silvery) pubescent throughout, trichomes several-rayed, rays furcate or bifurcate, (relatively massive, smooth to few-tubercled). | Perennials; (forming loose mats); caudex (buried), branched; densely pubescent, trichomes (short-stalked), 4–6-rayed, rays distinct, bifurcate, (rough-tuberculate). | ||||||||
Stems | several from base, usually decumbent to ascending, (arising laterally, unbranched, coarse), 1–2 dm. |
few to several from base, prostrate to erect or spreading, 0.05–0.5(–1.2) dm. |
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Basal leaves | blade pandurate or obovate, 3–6 cm, margins usually deeply and broadly incised, rarely subentire, (apex obtuse). |
(petiole and blade differentiated or not); blade spatulate to elliptic or linear or narrowly oblanceolate, 0.5–1.5(–2.5) cm, margins entire. |
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Cauline leaves | blade oblanceolate to spatulate, similar to basal, (3–6 mm wide), margins entire, (apex often somewhat acute). |
similar to basal, smaller. |
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Racemes | congested, (elongated in fruit). |
dense. |
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Flowers | sepals oblong, 6–8 mm; petals spatulate, to 10 mm. |
sepals narrowly lanceolate to lanceolate, 2.8–6 mm; petals (pale to deep yellow), narrowly lanceolate to oblanceolate, 5–9 mm, (claw undifferentiated from blade). |
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Fruiting pedicels | (usually curving upward, sigmoid), 6–10 mm. |
(ascending, straight or slightly curved), 2–6 mm. |
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Fruits | didymous, irregular in shape, somewhat angular, inflated, 5–7 × 6–8 mm, (papery, often rigid, base obtuse or truncate, apical sinus broad, open and deep); valves (retaining seeds after dehiscence), pubescent, trichomes spreading, loose; replum oblong, often constricted, as wide as or wider than fruit, apex obtuse; ovules 4 per ovary; style 5–7 mm. |
(sessile or substipitate), globose or subglobose to obovoid, not or slightly inflated, 3–6 mm, (firm, apex acute); valves (reddish in age, not retaining seeds after dehiscence), glabrous throughout; replum as wide as or wider than fruit; ovules 4–8 per ovary; style 1.7–6 mm. |
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Seeds | flattened. |
flattened. |
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2n | = 8, 16. |
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Physaria vitulifera |
Physaria hitchcockii |
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Phenology | Flowering Apr–Jun. | |||||||||
Habitat | Rocky hillsides, dry banks, gravel and sand, granitic slopes, soil scree, red shale | |||||||||
Elevation | 1600-3000 m (5200-9800 ft) | |||||||||
Distribution |
CO
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NV; UT |
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Discussion | Subspecies 3 (3 in the flora). The taxonomic treatment of Physaria hitchcockii has varied widely over the years. Molecular study (pers. obs.) has shown no direct relationship to P. tumulosa; morphologically, though, P. navajoensis and P. tumulosa appear closely related. Infraspecific taxonomy is based on the presence of a discernable petiole and whether or not the caudex is elastically elongated. The subspecies recognized here are usually geographically coherent, except that collections from the Table Cliff Plateau are more similar to subsp. hitchcockii, disjunct in Nevada, than they are to the very nearly sympatric subsp. rubicundula. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
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Key |
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Source | FNA vol. 7, p. 665. | FNA vol. 7, p. 643. | ||||||||
Parent taxa | Brassicaceae > tribe Physarieae > Physaria | Brassicaceae > tribe Physarieae > Physaria | ||||||||
Sibling taxa | ||||||||||
Subordinate taxa | ||||||||||
Synonyms | Lesquerella hitchcockii | |||||||||
Name authority | Rydberg: Bull. Torrey Bot. Club 28: 278. (1901) | (Munz) O’Kane & Al-Shehbaz: Novon 12: 324. (2002) | ||||||||
Web links |