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roundtip twinpod

Goodding's bladderpod

Habit Perennials; caudex simple or branched, (cespitose); (silvery) pubescent throughout, trichomes several-rayed, rays furcate or bifurcate, (relatively massive, smooth to few-tubercled). Annuals or biennials; without caudex, (cespitose); densely pubescent, trichomes (sessile or short-stalked), few-rayed, rays (ascending or erect), simple or infrequently furcate near base, (long and slender, sometimes with U-shaped notch on one side, smooth or finely tuberculate).
Stems

several from base, usually decumbent to ascending, (arising laterally, unbranched, coarse), 1–2 dm.

several from base, erect (and stout) or outer ones decumbent, (unbranched or branched, stiff and densely foliate, sterile leaf-bearing branches sometimes present), to 4 dm.

Basal leaves

blade pandurate or obovate, 3–6 cm, margins usually deeply and broadly incised, rarely subentire, (apex obtuse).

blade obovate or elliptic, to ca. 3 cm, margins sinuate or shallowly dentate.

Cauline leaves

blade oblanceolate to spatulate, similar to basal, (3–6 mm wide), margins entire, (apex often somewhat acute).

(proximal usually shortly petiolate, distal sessile);

blade obovate to broadly elliptic, 1–3 cm, margins sinuate or shallowly toothed.

Racemes

congested, (elongated in fruit).

dense, compact, (elongated in fruit).

Flowers

sepals oblong, 6–8 mm;

petals spatulate, to 10 mm.

sepals elliptic or narrowly elliptic or oblong, (3.8–)4.5–5.5 mm, (lateral pair cucullate, very convex, median pair tapering to base, thickened apically, cucullate, often slightly keeled);

petals cuneate, 6.5–8 mm, (slightly expanded at base, margins sinuate, apex retuse or entire).

Fruiting pedicels

(usually curving upward, sigmoid), 6–10 mm.

(recurved, curved or sigmoid), somewhat expanded apically.

Fruits

didymous, irregular in shape, somewhat angular, inflated, 5–7 × 6–8 mm, (papery, often rigid, base obtuse or truncate, apical sinus broad, open and deep);

valves (retaining seeds after dehiscence), pubescent, trichomes spreading, loose;

replum oblong, often constricted, as wide as or wider than fruit, apex obtuse;

ovules 4 per ovary;

style 5–7 mm.

(sessile or substipitate), oblong or broadly elliptic, compressed (latiseptate), 5–8 mm;

valves pubescent, trichomes spreading, sparsely pubescent inside;

ovules 4–6 per ovary;

style 3–5 mm.

Seeds

flattened.

flattened.

2n

= 8, 16.

Physaria vitulifera

Physaria gooddingii

Phenology Flowering Apr–Jun. Flowering Jun–Sep.
Habitat Rocky hillsides, dry banks, gravel and sand, granitic slopes, soil scree, red shale Mountainous areas, open areas in pinyon-juniper and ponderosa pine forests
Elevation 1600-3000 m (5200-9800 ft) 1800-2300 m (5900-7500 ft)
Distribution
from FNA
CO
[WildflowerSearch map]
[BONAP county map]
from FNA
AZ; NM
[BONAP county map]
Discussion

Physaria gooddingii (found in the mountains of Catron, Sierra, and western Socorro counties, New Mexico, and in Greenlee County, Arizona) is similar to 9. P. aurea (found farther east), but differs in having trichomes with ascending or erect rays (rather than appressed) and fruits that are strongly latiseptate (rather than not, or very little, compressed), a state that is infrequent in the genus.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 7, p. 665. FNA vol. 7, p. 639.
Parent taxa Brassicaceae > tribe Physarieae > Physaria Brassicaceae > tribe Physarieae > Physaria
Sibling taxa
P. acutifolia, P. alpestris, P. alpina, P. angustifolia, P. arctica, P. arenosa, P. argyraea, P. arizonica, P. aurea, P. bellii, P. brassicoides, P. calcicola, P. calderi, P. carinata, P. chambersii, P. cinerea, P. condensata, P. congesta, P. cordiformis, P. curvipes, P. densiflora, P. didymocarpa, P. dornii, P. douglasii, P. eburniflora, P. engelmannii, P. eriocarpa, P. fendleri, P. filiformis, P. floribunda, P. fremontii, P. garrettii, P. geyeri, P. globosa, P. gooddingii, P. gordonii, P. gracilis, P. grahamii, P. hemiphysaria, P. hitchcockii, P. humilis, P. integrifolia, P. intermedia, P. kingii, P. klausii, P. lata, P. lepidota, P. lesicii, P. lindheimeri, P. ludoviciana, P. macrocarpa, P. mcvaughiana, P. montana, P. multiceps, P. navajoensis, P. nelsonii, P. newberryi, P. obcordata, P. obdeltata, P. occidentalis, P. oregona, P. ovalifolia, P. pachyphylla, P. pallida, P. parviflora, P. parvula, P. pendula, P. pinetorum, P. prostrata, P. pruinosa, P. pulvinata, P. purpurea, P. pycnantha, P. rectipes, P. recurvata, P. reediana, P. rollinsii, P. saximontana, P. scrotiformis, P. sessilis, P. spatulata, P. subumbellata, P. tenella, P. thamnophila, P. tumulosa, P. valida, P. vicina
P. acutifolia, P. alpestris, P. alpina, P. angustifolia, P. arctica, P. arenosa, P. argyraea, P. arizonica, P. aurea, P. bellii, P. brassicoides, P. calcicola, P. calderi, P. carinata, P. chambersii, P. cinerea, P. condensata, P. congesta, P. cordiformis, P. curvipes, P. densiflora, P. didymocarpa, P. dornii, P. douglasii, P. eburniflora, P. engelmannii, P. eriocarpa, P. fendleri, P. filiformis, P. floribunda, P. fremontii, P. garrettii, P. geyeri, P. globosa, P. gordonii, P. gracilis, P. grahamii, P. hemiphysaria, P. hitchcockii, P. humilis, P. integrifolia, P. intermedia, P. kingii, P. klausii, P. lata, P. lepidota, P. lesicii, P. lindheimeri, P. ludoviciana, P. macrocarpa, P. mcvaughiana, P. montana, P. multiceps, P. navajoensis, P. nelsonii, P. newberryi, P. obcordata, P. obdeltata, P. occidentalis, P. oregona, P. ovalifolia, P. pachyphylla, P. pallida, P. parviflora, P. parvula, P. pendula, P. pinetorum, P. prostrata, P. pruinosa, P. pulvinata, P. purpurea, P. pycnantha, P. rectipes, P. recurvata, P. reediana, P. rollinsii, P. saximontana, P. scrotiformis, P. sessilis, P. spatulata, P. subumbellata, P. tenella, P. thamnophila, P. tumulosa, P. valida, P. vicina, P. vitulifera
Synonyms Lesquerella gooddingii
Name authority Rydberg: Bull. Torrey Bot. Club 28: 278. (1901) (Rollins & E. A. Shaw) O’Kane & Al-Shehbaz: Novon 12: 323. (2002)
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