Physaria vitulifera |
Physaria douglasii |
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roundtip twinpod |
Columbia bladder-pod, Douglas' bladder-pod |
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Habit | Perennials; caudex simple or branched, (cespitose); (silvery) pubescent throughout, trichomes several-rayed, rays furcate or bifurcate, (relatively massive, smooth to few-tubercled). | Perennials; caudex simple; densely pubescent, trichomes (sessile or nearly so), 4–6(–10)-rayed, rays usually furcate near base, rarely bifurcate, (umbonate, tuberculate throughout). | ||||
Stems | several from base, usually decumbent to ascending, (arising laterally, unbranched, coarse), 1–2 dm. |
simple from base, erect, (usually unbranched), to 4.5 dm. |
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Basal leaves | blade pandurate or obovate, 3–6 cm, margins usually deeply and broadly incised, rarely subentire, (apex obtuse). |
blade suborbicular to elliptic, 2–9.5(–11.5) cm, margins entire, sinuate, coarsely dentate, or almost lyrate-pinnatifid. |
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Cauline leaves | blade oblanceolate to spatulate, similar to basal, (3–6 mm wide), margins entire, (apex often somewhat acute). |
similar to basal, blade narrowly linear or, sometimes, orbicular. |
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Racemes | congested, (elongated in fruit). |
loose (lax). |
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Flowers | sepals oblong, 6–8 mm; petals spatulate, to 10 mm. |
sepals elliptic or ovate, (2–)3.5–7.5 mm, (cucullate); petals 6–11 mm. |
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Fruiting pedicels | (usually curving upward, sigmoid), 6–10 mm. |
(recurved, straight, curved, or sigmoid), 6–20 mm. |
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Fruits | didymous, irregular in shape, somewhat angular, inflated, 5–7 × 6–8 mm, (papery, often rigid, base obtuse or truncate, apical sinus broad, open and deep); valves (retaining seeds after dehiscence), pubescent, trichomes spreading, loose; replum oblong, often constricted, as wide as or wider than fruit, apex obtuse; ovules 4 per ovary; style 5–7 mm. |
obovoid to subglobose, not inflated (not angustiseptate), 3–6 mm; valves sparsely pubescent, sometimes glabrous inside, trichomes sessile or stalked; ovules 4(–8) per ovary; style (1.6–)3–6 mm. |
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Seeds | flattened. |
flattened. |
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2n | = 8, 16. |
= 10, 30. |
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Physaria vitulifera |
Physaria douglasii |
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Phenology | Flowering Apr–Jun. | |||||
Habitat | Rocky hillsides, dry banks, gravel and sand, granitic slopes, soil scree, red shale | |||||
Elevation | 1600-3000 m (5200-9800 ft) | |||||
Distribution |
CO
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ID; MT; OR; WA; BC
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Discussion | Subspecies 2 (2 in the flora). (Discussion copyrighted by Flora of North America; reprinted with permission.) |
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Key |
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Source | FNA vol. 7, p. 665. | FNA vol. 7, p. 634. | ||||
Parent taxa | Brassicaceae > tribe Physarieae > Physaria | Brassicaceae > tribe Physarieae > Physaria | ||||
Sibling taxa | ||||||
Subordinate taxa | ||||||
Synonyms | Lesquerella douglasii | |||||
Name authority | Rydberg: Bull. Torrey Bot. Club 28: 278. (1901) | (S. Watson) O’Kane & Al-Shehbaz: Novon 12: 322. (2002) | ||||
Web links |
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